生物全息数据库系统

基本信息

学名(拉丁名)	Viburnum
中文名	荚蒾属
命名人	Linn.
中文科名	忍冬科
类型	genus
拉丁科名	Caprifoliaceae

学名(拉丁名)	Viburnum
中文名	荚蒾属
中文科名	忍冬科
类型	genus
拉丁科名	Caprifoliaceae

学名(拉丁名)	Viburnum
类型	genus
拉丁科名	Adoxaceae

学名(拉丁名)	Viburnum
中文名	荚蒾属
类型	genus
拉丁科名	Adoxaceae

学名(拉丁名)	Viburnum
中文名	荚蒾属
中文科名	忍冬科
类型	genus
拉丁科名	Caprifoliaceae

学名(拉丁名)	Viburnum
发表年份	1753(年)
类型	genus
拉丁科名	Adoxaceae

学名(拉丁名)	Viburnum
类型	genus

名称状态

学名	状态	来源
Viburnum	接受名	The Plant List
Viburnum	接受名	云南植物志
Viburnum	接受名	中国植物志
Viburnum	接受名	Flora of China

下级分类单元

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foc

1	冬芽裸露；植物体被簇状毛而无鳞片；果实成熟时由红色转为黑色。 (2)
1	冬芽有1-2对（很少3对或多对）鳞片1）；如为裸露，则芽、幼枝、叶下面、花序、萼、花冠及果实均被鳞片状毛。 (19)
2	花序有总梗；果核有2条背沟和 (1-) 3条腹沟，或有时背沟退化而不明显；胚乳坚实（组1. 裸牙组 Sect. Viburnum）。 (3)	裸芽组 Sect. Viburnum
2	花序无总梗；果核有1条背沟和1条深腹沟；胚乳深嚼烂状（组2. 合轴组 Sect. Pseudotinus C. B. Clarke） (18)	合轴组 Sect. Pseudotinus
3	叶临冬凋落，通常边缘有齿。 (4)
3	叶大多常绿，全缘或有时具不明显的疏浅齿，侧脉通常近叶缘时互相网结而非直达齿端。 (15)
4	叶的侧脉近叶缘时虽分枝，但直达齿端而非互相网结，或至少大部分如此。 (5)
4	叶的侧脉近叶缘时互相网结而非直达齿端，或至少大部分如此。 (9)
5	萼筒无毛；花冠裂片比筒短。	黄栌叶荚蒾 Viburnum cotinifolium
5	萼筒被簇状毛；花冠裂片比筒长或几相等。 (6)
6	叶矩圆状披针形，长度为阔度的3倍以上；雄蕊不高出花冠；花冠直径7-9毫米，裂片比筒长。	醉鱼草状荚蒾 Viburnum buddleifolium
6	叶卵形、宽卵形至卵状矩圆形，有时倒卵形至倒卵状矩圆形，长度为阔度的2倍或不到；雄蕊稍高出花冠；花冠直径5-7毫米。 (7)
7	叶的侧脉5-6对；花序直径约4厘米。	圆叶荚蒾 Viburnum glomeratum subsp. rotundifolium
7	叶的侧脉6-11对；花序直径3-10厘米。 (8)
8	叶卵形至宽卵形，通常长不超过10厘米（很少达15厘米）；花序直径3-6厘米；果核长5-7 (-9) 毫米。	聚花荚蒾 Viburnum glomeratum
8	叶大，卵状矩圆形至卵形，长10-19厘米；花序直径8-10厘米；果核长9-11毫米。	壮大荚蒾 Viburnum glomeratum subsp. magnificum
9	花序有大型的不孕花。 (10)
9	花序全由两性花组成，无大型的不孕花。 (11)
10	花序全部由大型的不孕花组成。	绣球荚蒾 Viburnum macrocephalum
10	花序仅周围有大型的不孕花。	琼花 Viburnum macrocephalum form. keteleeri
11	花冠辐状，筒比裂片短。 (12)
11	花冠筒状钟形，筒远比裂片长。 (14)
12	二年生小枝黄白色；叶顶端通常尖，稀稍钝，基部两侧常不对等；花大部生于花序的第二级辐射枝上；果核扁，有2条背沟。	修枝荚蒾 Viburnum burejaeticum
12	二年生小枝灰褐色；叶顶端钝或圆形，稀稍尖；花大部生于花序的第三至第四级辐射枝上；果核背部凸起而无沟，长6-8毫米。 (13)
13	萼筒无毛。	陕西荚蒾 Viburnum schensianum
13	萼筒被簇状短毛。	浙江荚蒾 Viburnum schensianum subsp. chekiangense
14	叶宽卵形、椭圆形至近圆形，长2.5-6厘米，顶端尖或钝形；叶柄长4-10毫米；花生于花序的第一级（稀第二级）辐射枝上；花冠黄白色。	蒙古荚蒾 Viburnum mongolicum
14	叶卵状披针形或卵状矩圆形，长7-15厘米，顶端长渐尖；叶柄较长，长1-4厘米；花生于花序的第二至第四级辐射枝上；花冠外面紫红色。	壶花荚蒾 Viburnum urceolatum
15	萼筒无毛；叶长2-6厘米（烟管荚蒾 V. utile Hemsl. 有时达8. 5厘米），上面小脉不凹陷。 (16)
15	萼筒多少被簇状毛；叶长5-25厘米。 (17)
16	花冠钟状漏斗形，裂片短于筒；老叶下面的簇状毛均匀而不完全掩盖整个表面。	密花荚蒾 Viburnum congestum
16	花冠辐状，裂片与筒等长或略较长；老叶下面的簇状毛覆盖整个表面。	烟管荚蒾 Viburnum utile
17	叶披针状矩圆形至狭矩圆形，通常长515厘米，宽1.5-4.5厘米，老时厚纸质，上面侧脉或有时连同小脉略凹陷，但不为极度皱纹状；叶柄长1-2厘米；花冠外面疏被簇状毛。	金佛山荚蒾 Viburnum chinshanense
17	叶卵状披针形至卵状矩圆形，通常长8-25厘米，宽2.5-8厘米，革质，上面各脉均深凹陷，呈现极度的皱纹状；叶柄长1.5-4厘米；花冠外面几无毛。	皱叶荚蒾 Viburnum rhytidophyllum
18	花序无大型的不孕花。	显脉荚蒾 Viburnum nervosum
18	花序周围有大型的不孕花。	合轴荚蒾 Viburnum sympodiale
19	果核圆形、卵圆形或椭圆形，有1条极细的线状浅腹沟或无沟，决不带压扁状；花序复伞形式；果实成熟时蓝黑色或由蓝色转为黑色；叶常绿，无毛或近无毛（组3. 球核组 Sect. Tinus (Borkh.) Maxim.） (20)	球核组 Sect. Tinus (Borkh.) Maxim.()
19	果核不如上述；如为椭圆形则果核具1上宽下窄的深腹沟，或花序不如上述；果实成熟时红色，或由红色转为黑色或酱黑色，少有黄色。 (26)
20	叶具羽状脉。 (21)
20	叶具三出脉或离基三出脉。 (22)
21	叶圆卵形至卵状披针形或菱状椭圆形，长3-10厘米，顶端钝而有小凸尖，边缘常有不规则小尖齿。	蓝黑果荚蒾 Viburnum atrocyaneum
21	叶椭圆状矩圆形、宽卵形或倒卵形，较小，长0.8-6厘米，顶端钝至圆形或微凹缺而有小凸尖，全缘或有不规则锯齿。	毛枝荚蒾 Viburnum atrocyaneum subsp. harryanum
22	叶顶端钝至圆形，长2-6 (-7.5) 厘米，全缘；总花梗长约1厘米。	三脉叶荚蒾 Viburnum triplinerve
22	叶顶端锐尖至渐尖；总花梗长1. 5-4 (-5) 厘米。 (23)
23	花冠暗红色；老叶厚革质，上面常因小脉深凹陷而呈明显的皱纹状。	川西荚蒾 Viburnum davidii
23	花冠绿白色或淡黄绿色；老叶革质，有时上面小脉略凹陷，但不为明显的皱纹状。 (24)
24	花序大而松散，直径6-15厘米；叶长6-13 (-18) 厘米，全缘或近顶端偶有少数锯齿。	樟叶荚蒾 Viburnum cinnamomifolium
24	花序小，直径4-5厘米，无毛；叶长3-9 (-11) 厘米，边缘通常有锯齿。 (25)
25	叶卵形至卵状披针形或椭圆形至椭圆状矩圆形，基部近圆形或阔楔形至楔形。	球核荚蒾 Viburnum propinquum
25	叶条状披针形或倒披针形，基部楔形。	狭叶球核荚蒾 Viburnum propinquum var. mairei
26	冬芽有1-2对分离的鳞片；叶柄顶端或叶片基部无腺体。 (27)
26	冬芽为2对合生的鳞片所包围；叶3 (2-4) 裂；叶柄顶端或叶片基部有2-4个明显的腺体（组9. 裂叶组 Sect. Opulus DC.）。 (106)	裂叶组 Sect. Opulus
27	叶掌状3-5裂，具掌状脉。	甘肃荚蒾 Viburnum kansuense
27	叶不分裂或规则至不规则2-3浅裂，大多具羽状脉，有时基部一对侧脉近似三出脉或离基三出脉。 (28)
28	花序复伞形或伞形式，有大型的不孕花；果核腹面有1上宽下窄的沟，沟上端及背面下半部中央各有1明显隆起的脊（组5. 蝶花组 Sect. Pseudo pulus (Dipp.) Rehd.） (29)	蝶花组 Sect. Pseudopulus
28	花序种种，不具大型不孕花；果核通常不如上述。 (31)
29	叶有5-7 (-9) 对侧脉，两面长方形格纹不明显；总花梗的第一级辐射枝通常5条。	蝶花荚蒾 Viburnum hanceanum
29	叶有10对以上侧脉，两面有明显的长方形格纹；总花梗的第一级辐射枝6-8条。 (30)
30	花序全部由大型的不孕花组成。	粉团 Viburnum plicatum
30	仅花序周围有4-6朵大型的不孕花。	蝴蝶戏珠花 Viburnum plicatum var. tomentosum
31	花序为由穗状或总状花序组成的圆锥花序，或因圆锥花序的主轴缩短而近似伞房式，很少花序紧缩成近簇状；果核通常浑圆或稍扁，具1上宽下窄的深腹沟（组4. 圆锥组 Sect. Thyrsosms (Rafin.) Rehd.） (32)	圆锥组 Sect. Thyrsosma
31	花序复伞形式或稀可为由伞形花序组成的尖塔形圆锥花序；果核通常扁，有浅的背、腹沟，有时沟退化而不明显，很少无沟或在腹面深陷如杓状。 (63)
32	花冠漏斗形或高脚蝶形，很少辐状钟形，裂片短于筒。 (33)
32	花冠辐状，裂片长于筒。 (56)
33	雄蕊着生于花冠筒内的不同高度；花先于叶或与叶同时开放；叶纸质。 (34)
33	雄蕊着生于花冠筒顶端；花于叶后（极少与叶同时）开放；叶纸质至革质。 (35)
34	花序圆锥式，生于具幼叶的短枝之顶；苞片近无毛。	香荚蒾 Viburnum farreri
34	花序紧缩成近簇状，生于无叶的短枝之顶；苞片初时密被银白色绢毛。	大花荚蒾 Viburnum grandiflorum
35	叶纸质，下面脉腋有趾蹼状小孔；聚伞花序长3-4厘米。	短筒荚蒾 Viburnum brevitubum
35	叶下面脉腋无趾蹼状小孔，如有小孔则叶革质而非纸质；花序通常长6厘米以上。 (36)
36	叶的侧脉大部分直达齿端；叶纸质。 (37)
36	叶的侧脉大部分在近叶缘时互相网结；叶纸质、厚纸质至革质。 (41)
37	花序极小，长约1.5厘米；花生于序轴的第一级辐射枝上；叶长2-3厘米，边缘有钝锯齿。	小红荚蒾 Viburnum erubescens var. parvum
37	花序长2.5厘米以上，花生于序轴的第一至第三级辐射枝上；叶长4.5厘米以上，边缘有锐锯齿。 (38)
38	花冠辐状钟形，筒长约3毫米；雄蕊明显高出花冠筒。	瑞丽荚蒾 Viburnum shweliense
38	花冠高脚碟形，筒长5-6 (-8) 毫米；雄蕊稍高出花冠筒。 (39)
39	花药堇紫色。	紫药红荚蒾 Viburnum erubescens var. prattii
39	花药黄白色。 (40)
40	花无梗或有短梗；叶通常椭圆形至狭矩圆形，很少卵状心形或略带倒卵形。	红荚蒾 Viburnum erubescens
40	花序中之一部分花有细长梗；叶多少呈倒卵形，基部带圆形。	细梗红荚蒾 Viburnum erubescens var. gracilipes
41	圆锥花序较小，通常直径在5厘米以下。 (42)
41	圆锥花序较大，通常直径在5厘米以上，具多数稠密的花少至少部分花生于序轴的第三或第四级辐射枝上。 (52)
42	叶下面无红褐色微细腺点。 (43)
42	叶下面在放大镜下可见红褐色微细腺点。 (50)
43	叶边缘有尖或钝的锯齿，齿顶不向内或向前弯。 (44)
43	叶边缘有尖锯齿，齿顶通常向内或向前弯。 (49)
44	苞片绿色，叶质，矩圆形或条形；花小，花冠筒长约3. 5毫米，裂片之较余者稍大。	峨眉荚蒾 Viburnum omeiense
44	苞片膜质，非绿色叶质；花较大，花冠筒长4-10毫米，各裂片大小大致相等。 (45)
45	果核有1条深陷的封闭式管形腹沟，横切面呈不规则六角形。	台东荚蒾 Viburnum taitoense
45	果核有1条宽广的腹沟，横切面呈扁圆形。 (46)
46	叶上面侧脉和小脉凹陷。	凹脉肉叶荚蒾 Viburnum chingii var. impressinervium
46	叶上面侧脉和小脉不凹陷。 (47)
47	叶较狭，矩圆状椭圆形或倒披针状矩圆形，宽1.5-2.5厘米；总花梗长1-2厘米；花冠淡红色。	肉叶荚蒾 Viburnum chingii var. carnosulum
47	叶卵状椭圆形至倒卵形或倒卵状矩圆形，很少卵状披针形，宽2-4. 5厘米；总花梗长 (2-) 3.5-6.5厘米；花冠白色。 (48)
48	总花梗长 (2-) 3.5-4.5 (-6) 厘米；叶边缘有钝或尖的锯齿，齿开展或否。	漾濞荚蒾 Viburnum chingii
48	总花梗长4-6.5厘米，颇纤细；叶边缘离基1/3以上有开展的小尖齿。	细梗漾濞荚蒾 Viburnum chingii var. tenuipes
49	叶亚革质至革质，少有厚纸质，上面中脉明显凸起；花药紫红色；果实长6-7毫米；总花梗长 (1.2-) 2.5-7厘米。	少花荚蒾 Viburnum oliganthum
49	叶纸质，上面中脉不明显凸起；花药黄白色；果实长8-10毫米；总花梗长 (3.5-) 6-9厘米。	长梗荚蒾 Viburnum longipedunculatum
50	叶边缘有锐锯齿；总花梗直立。	肉叶荚蒾 Viburnum chingii var. carnosulum
50	叶边缘有钝锯齿；总花梗弯垂。 (51)
51	叶长3-6厘米，边缘每5毫米内约有1-2个锯齿。	亚高山荚蒾 Viburnum subalpinum
51	叶较小，长2.5-4.5厘米，边缘每5毫米内约有3-4个锯齿。	边沿荚蒾 Viburnum subalpinum var. limitaneum
52	果核略扁，横切面呈扁圆形；叶纸质或厚纸质，边缘有较规则的尖或稍尖的锯齿。 (53)
52	果核浑圆，横切面呈近圆形；叶革质，近全缘或有波状钝锯齿或不规则粗钝牙齿。 (55)
53	圆锥花序尖塔形；叶有7-8对侧脉；二年生小枝淡灰褐色。	横脉荚蒾 Viburnum trabeculosum
53	圆锥花序因序轴缩短而呈复伞房式；叶有4-6对侧脉；二年生小枝紫褐色。 (54)
54	叶矩圆形或倒卵状矩圆形，下面无毛或仅中脉和侧脉有少数簇状短毛。	滇缅荚蒾 Viburnum burmanicum
54	叶倒卵形至倒披针形或卵状椭圆形，下面全面被簇状短毛。	墨脱荚蒾 Viburnum burmanicum var. motoense
55	苞片长不到1厘米，宽不及2毫米，或不存在；果核长6-7毫米，通常倒卵圆形至倒卵状椭圆形。	日本珊瑚树 Viburnum odoratissimum var. awabuki
55	苞片大，长可达2厘米，宽达5毫米；果核长约9毫米，卵状椭圆形。	云南珊瑚树 Viburnum odoratissimum var. sessiliflorum
56	叶纸质，下面在放大镜下可见红褐色微细腺点；幼枝、叶柄和花序均被黄褐色、通常呈绒状的簇状柔毛。	云南荚蒾 Viburnum yunnanense
56	叶厚纸质至革质，下面无红褐色腺点。 (57)
57	圆锥花序尖塔形；如因花序轴稍缩短而花序近似伞房式，则叶下面脉腋有趾蹼状小孔。 (58)
57	圆锥花序因序轴不充分伸长而呈圆顶的，外观近似伞房式；如为短圆锥式，则叶下面脉腋无小孔。 (62)
58	叶的侧脉至少一部分直达齿端；花序无毛或近无毛。	巴东荚蒾 Viburnum henryi
58	叶的侧脉近叶缘时弯拱而互相网结，不直达齿端。 (59)
59	叶革质；果核卵圆形、卵状椭圆形至长卵圆形。 (60)
59	叶厚纸质；果核略为压扁状。 (61)
60	萼和花冠均无毛；果核卵圆形或卵状椭圆形，顶端常多少骤然收缩而带圆形，因而有肩。	珊瑚树 Viburnum odoratissimum
60	萼和花冠或至少萼外面被簇状短毛；果核卵圆形或长卵圆形，顶端常渐尖而无肩，未熟果实常疏生簇状毛。	短序荚蒾 Viburnum brachybotryum
61	叶有5-6对侧脉，边缘有尖锯齿。	腾越荚蒾 Viburnum tengyuehense
61	叶有6-10对侧脉，边缘有钝锯齿。	多脉腾越荚蒾 Viburnum tengyuehense var. polyneurum
62	小枝黄白色；叶矩圆形或矩圆状披针形，长6-13厘米。	伞房荚蒾 Viburnum corymbiflorum
62	小枝淡灰褐色；叶椭圆形至倒卵形，长5-9.5厘米。	苹果叶荚蒾 Viburnum corymbiflorum subsp. malifolium
63	冬芽有1对鳞片，极少裸露。 (64)
63	冬芽有2对鳞片（组8. 齿叶组 Sect. Odontotinus Rehd.）。 (74)	齿叶组 Sect. Odontotinus
64	叶的侧脉在近叶缘时弯拱而互相网结，不直达齿端；花序生于有1至多对叶的小枝之顶（组7. 鳞腺组 Sect. Megalotinus (Maxim.) Rehd.） (65)	大叶组 Sect. Megalotinus
64	叶的侧脉全部或至少部分直达齿端；花序生于有1对叶的侧生短枝之顶（组6. 侧花组 Sect. Platyphylla Hsu） (73)	侧花组 Sect. Platyphylla
65	冬芽裸露；芽、幼枝、叶下面、花序和花冠外面均被铁锈色、圆形鳞片状毛。 (66)
65	冬芽有1对鳞片；植物体无上迷鳞片状毛。 (67)
66	花冠直径约6毫米；果实长8-10毫米，宽6-8毫米。	鳞斑荚蒾 Viburnum punctatum
66	花冠直径约8毫米；果实长14-15 (-18) 毫米，宽约10毫米。	大果鳞斑荚蒾 Viburnum punctatum var. lepidotulum
67	花冠钟状，裂片短而直立；叶下面有带红色或黄色腺点（腺点有时扁化而类似鳞片）。	水红木 Viburnum cylindricum
67	花冠辐状。 (68)
68	叶全缘或有时顶端有少数大牙齿，基部中脉两侧常有大形腺斑；雄蕊远高出于花冠，花丝在蕾中褶叠。 (69)
68	叶边缘除基部外有牙齿或锯齿，基部两侧无腺斑；雄蕊稍长于花冠，花丝在蕾中不褶叠。 (72)
69	叶通常3枚轮生；托叶2枚；花序无或近于无总花梗。	三叶荚蒾 Viburnum ternatum
69	叶对生；托叶不存在或早落；花序有总花梗。 (70)
70	叶下面被厚绒状簇状毛；萼筒被长柔毛状簇状毛。	厚绒荚蒾 Viburnum inopinatum
70	叶下面被短柔毛或仅脉上有毛；萼筒无毛。 (71)
71	叶上面仅中脉疏被短柔毛，无凸起细点，下面无腺点。	光果荚蒾 Viburnum leiocarpum
71	叶上面全被簇状或叉状短柔毛，有凸起细点，下面有腺点。	斑点光果荚蒾 Viburnum leiocarpum var. punctatum
72	除小枝和叶下面初时疏被簇状短毛，后告脱落外，全体近于无毛；花序通常复伞形式。	淡黄荚蒾 Viburnum lutescens
72	小枝、叶下面、叶柄、总花梗、花梗和萼均被由宿存的黄褐色簇状毛组成的绒毛；花序常为由2-4层伞形花序组成的尖塔形圆锥花序。	锥序荚蒾 Viburnum pyramidatum
73	幼枝、叶柄和花序均被由带黄褐色簇状毛组成的绒毛；总花梗的第一级辐射枝6-7条。	广叶荚蒾 Viburnum amplifolium
73	全株无毛；总花梗的第一级辐射枝5条。	侧花荚蒾 Viburnum laterale
74	叶的侧脉2-4对，基部1对作离基或近离基三出脉状；如侧脉5-6对，则叶革质或亚革质；或叶纸质或厚纸质而下面在放大镜下同时可见具金黄色和红褐色至黑褐色两种腺点。 (75)
74	叶的侧脉5对以上，羽状，很少类似离基三出脉；叶纸质、厚纸质或薄革质，下面无腺点或有颜色纯一的腺点。 (85)
75	幼枝四方形。 (76)
75	幼枝圆柱状，纵有棱角亦不为四方形。 (80)
76	叶下面有金黄色和红褐色至黑褐色两种腺点，干后上面通常不变黑色。 (77)
76	叶下面有黑色或栗褐色腺点（在放大镜下可见），干后上面变黑色。 (78)
77	幼枝和叶柄无毛或近无毛；叶柄和花序疏被黄褐色短伏毛或近无毛；叶除下面脉腋有时具簇聚毛外，无毛。	金腺荚蒾 Viburnum chunii
77	幼枝和叶柄均密被短伏毛；叶上面近边缘处疏被短伏毛，下面中脉和侧脉疏生短毛。	毛枝金腺荚蒾 Viburnum chunii var. piliferum
78	萼筒被簇状毛；叶亚革质。	海南荚蒾 Viburnum hainanense
78	萼筒无毛；叶革质。 (79)
79	幼枝、叶柄和花序无毛或散生少数簇状短毛；果核背面凸起，腹面明显凹陷，其形如杓，宽3-5毫米。	常绿荚蒾 Viburnum sempervirens
79	幼枝、叶柄和花序均密被簇状短毛；果核背面略凸起，腹面略呈鹅毛扇状弯拱而不明显凹陷，长约7毫米，宽约6毫米。	具毛常绿荚蒾 Viburnum sempervirens var. trichophorum
80	叶两面无毛，全缘，顶端急狭而长尾尖。	全叶荚蒾 Viburnum integrifolium
80	叶至少下面有簇状毛。 (81)
81	叶边缘有不规则、圆或钝的粗牙齿或缺刻，大多倒卵状椭圆形，长2-5厘米。	珍珠荚蒾 Viburnum foetidum var. ceanothoides
81	叶中部以上边缘常有少数浅齿或有时全缘。 (82)
82	叶通常长1-3厘米，下面脉腋无簇聚毛；果核有1条腹沟。	小叶荚蒾 Viburnum parvifolium
82	叶长3-6 (-10) 厘米，下面脉腋有簇聚毛；果核有3条腹沟。 (83)
83	枝不作披散状；小枝亦不甚伸长，不呈蜿蜓状；总花梗长 (0.5-) 2-5厘米。	臭荚蒾 Viburnum foetidum
83	枝披散；小枝伸长而往往蜿蜒状；总花梗极短或几不存，最长达2厘米。	直角荚蒾 Viburnum foetidum var. rectangulatum
84	果核背面凸起，腹面四周升高，中部凹陷，形状如杓；叶矩圆状披针形至披针形或条状披针形，长9-19厘米；幼枝四方形。 (85)
84	果核通常带扁形，有时可因两侧边缘向腹面反卷而纵向凹陷，但不为杓状；如多少带浅杓状则叶形不如上述。 (86)
85	幼枝、叶和叶柄均无毛；叶的侧脉约6对。	瑶山荚蒾 Viburnum squamulosum
85	幼枝、叶下面脉上（除叉状毛及单毛外）和叶柄均被簇状短毛；叶的侧脉7-14对。	披针叶荚蒾 Viburnum lancifolium
86	花冠外面无毛，极少蕾时有毛而花开后变秃净。 (87)
86	花冠外面被疏或密的簇状短毛。 (98)
87	花序或果序下垂；幼枝多少有棱角；芽及叶干后变黑色或浅灰黑色。 (88)
87	花序或果序不下垂。 (89)
88	果核两侧边缘不反卷，腹面扁平或略凹陷。	茶荚蒾 Viburnum setigerum
88	果核两侧边缘因向腹面反卷而明显地纵向凹陷。	沟核茶荚蒾 Viburnum setigerum var. sulcatum
89	总花梗长5.5-10 (-12.5) 厘米；叶有时顶端浅3裂或不规则分裂。	衡山荚蒾 Viburnum hengshanicum
89	总花梗通常长不超过5厘米；叶不分裂。 (90)
90	叶下面全面被带白色簇状毛。	卷毛荚蒾 Viburnum betulifolium var. flocculosum
90	叶下面仅中脉和侧脉被长伏毛或脉腋有集聚簇状毛。 (91)
91	叶下面在放大镜下可见透亮腺点。	浙皖荚蒾 Viburnum wrightii
91	叶下面无上述腺点。 (92)
92	总花梗的第一级辐射枝通常7出；花生于第 (3-) 4-5 (-6) 级辐射枝上；果实成熟时红色。	桦叶荚蒾 Viburnum betulifolium
92	总花梗的第一级辐射枝通常5出；花生于第2-4级辐射枝上。 (93)
93	果核多少呈浅杓状，腹面中央有1纵向隆起的脊；叶为宽狭程度不同的倒卵形、近圆形或宽椭圆形，稀菱状椭圆形，顶端通常突然收缩成短渐尖；叶柄有或无托叶；果实成熟时酱黑色。	黑果荚蒾 Viburnum melanocarpum
93	果核不为浅杓状；叶狭卵形、椭圆状卵形或菱状卵形，稀卵状披针形或倒卵形，顶端通常渐尖。 (94)
94	叶柄不具托叶。 (95)
94	叶柄有托叶。 (97)
95	萼筒无毛。	光萼荚蒾 Viburnum formosanum subsp. leiogynum
95	萼筒被簇状短毛。 (96)
96	幼枝无毛；叶柄有少数简单毛；花序薄被簇状短毛。	台中荚蒾 Viburnum formosanum
96	幼枝、叶柄和花序均密被黄褐色簇状毛，并夹杂简单长毛。	毛枝台中荚蒾 Viburnum formosanum subsp. formosanum var. pubigerum
97	叶不分裂，边缘有小尖齿。	宜昌荚蒾 Viburnum erosum
97	叶基部常浅2裂，边缘有粗牙齿或缺刻状牙齿。	裂叶宜昌荚蒾 Viburnum erosum var. taquetii
98	叶下面在放大镜下有黄色或近无色的透亮腺点，或有时腺点呈鳞片状，更或为不明显的暗色。 (99)
98	叶下面无腺点。 (101)
99	叶柄长仅2-3毫米；叶下面脉腋无集聚簇状毛；雄蕊约与花冠等长。	短柄荚蒾 Viburnum brevipes
99	叶柄长5毫米以上；叶下面脉腋有集聚簇状毛；雄蕊远高出花冠。 (100)
100	幼枝、叶柄和花序均密被刚毛状糙毛，很少幼枝有时无毛或近无毛；叶干后不变灰黑色或黑色。	荚蒾 Viburnum dilatatum
100	幼枝、叶柄和花序均密被黄褐色绒毛；叶干后至少上面变灰黑色至黑色。	庐山荚蒾 Viburnum dilatatum var. fulvotomentosum
101	叶上面有腺点；总花梗极短或几无，很少可长达1. 5厘米。	吕宋荚蒾 Viburnum luzonicum
101	叶上面无腺点；总花梗明显或罕有极短。 (102)
102	叶卵状披针形或卵状椭圆形，顶端常长渐尖，有8-12对侧脉。	粤赣荚蒾 Viburnum dalzielii
102	叶形通常不如上述，有5-9对侧脉。 (103)
103	花柱高出或略高出萼齿；花冠裂片比筒长；雄蕊与花冠等长或略高出；果核长6-7.5毫米。 (104)
103	花柱极短，比萼齿短；花冠裂片与筒几等长；雄蕊短于花冠；果核长4-6毫米。 (105)
104	叶顶端急尾尖，有7-9对侧脉；托叶存在或否；总花梗的第一级辐射枝5-7条。	长伞梗荚蒾 Viburnum longiradiatum
104	叶顶端短尖至短渐尖，有5-7 (-9) 对侧脉；托叶不存在；总花梗的第一级辐射枝通常5条。	南方荚蒾 Viburnum fordiae
105	当年小枝和幼叶下面被由簇状短毛组成的绒毛；萼筒和花冠外面密被簇状短毛。	西域荚蒾 Viburnum mullaha
105	年小枝近无毛；叶下面除脉腋有集聚簇状毛外至多脉上有毛；萼筒和花冠外面被极稀短毛。	少毛西域荚蒾 Viburnum mullaha var. glabrescens
106	花序无大型的不孕花；叶2-3 (-4) 裂；叶柄最长不超过2. 5厘米。	朝鲜荚蒾 Viburnum koreanum
106	花序周围有大型的不孕花；叶通常3裂或有时小枝上部同时存在不裂的叶；叶柄长2-4厘米。 (107)
107	树皮质薄而非木栓质；花药黄白色。	欧洲荚蒾 Viburnum opulus
107	树皮厚，木栓质；花药紫红色。 (108)
108	小枝、叶柄和总花梗均无毛；叶下面仅脉腋有集聚簇状毛，或有时脉上亦有少数柔毛。	鸡树条 Viburnum opulus var. calvescens
108	幼枝、叶下面和总花梗均被长柔毛。	毛叶鸡树条 Viburnum opulus var. calvescens

1	Inflorescence a panicle, sometimes densely clustered, rarely a corymblike panicle, rays opposite, decussate, first node of inflorescence with 2 rays; pyrenes with a deep ventral groove (if first node of panicle-like inflorescence with 4-6 rays and pyrenes with 2 deep dorsal grooves and 1 shallow ventral groove then see V. pyramidatum ) (4. V. sect. Solenotinus )	(2)
1 +	Inflorescence compound and umbel-like, rarely pyramidal panicle consisting of umbels; rays whorled, first node of inflorescence with 4-8 rays, (if first node of corymblike inflorescence with 2 rays and pyrenes with a deep ventral groove then see V. yunnanense and V. corymbiflorum ); pyrenes with 1 shallow ventral, or with both ventral and dorsal grooves	(19)
2 (1)	Corolla rotate, lobes as long as or longer than tube	(3)
2 (1) +	Corolla funnelform or hypocrateriform, very rarely rotate-campanulate, lobes shorter than tube	(8)
3 (2)	Leaves papery, abaxially with red-brown glandular dots; inflorescence a corymblike panicle. (38)	V. yunnanense
3 (2) +	Leaves thickly papery to leathery, abaxially without red-brown glandular dots; inflorescence a corymblike or pyramidal panicle	(4)
4 (3)	Panicle corymblike. (24)	V. corymbiflorum
4 (3) +	Panicle pyramidal	(5)
5 (4)	Lateral veins at least partly directly ending in teeth; inflorescence axes glabrous or subglabrous; calyx reddish. (28)	V. henryi
5 (4) +	Lateral veins arched and anastomosing near margin, not directly ending in teeth; inflorescence axes glabrous or stellate-pubescent; calyx green	(6)
6 (5)	Leaves thickly papery; pyrenes ca. 4 × 3.5 mm. (36)	V. tengyuehense
6 (5) +	Leaves leathery; pyrenes 7-8 × 4-5 mm	(7)
7 (6)	Calyx and corolla glabrous; flowers fragrant; pyrenes ovoid or ovoid-ellipsoid, apex rounded. (30)	V. odoratissimum
7 (6) +	Calyx and corolla, or at least calyx, stellate-pubescent outside; flowers not fragrant; pyrenes ovoid or narrowly ovoid, apex acuminate. (21)	V. brachybotryum
8 (2)	Stamens inserted at or above middle of corolla tube, generally at different heights; flowers opening before leaves; leaves papery	(9)
8 (2) +	Stamens inserted at apex of corolla tube; flowers opening after leaves; leaves papery to leathery	(10)
9 (8)	Leaves abaxially slightly pubescent on veins, veins 5-7-jugate, apex acute; inflorescence a panicle, at apices of short branchlets with young leaves; bracts ciliate, subglabrous. (26)	V. farreri
9 (8) +	Leaves abaxially densely pubescent, veins 6-10-jugate, apex acuminate; inflorescence densely clustered at apices of leafless short branchlets; bracts densely silvery sericeous initially. (27)	V. grandiflorum
10 (8)	Leaves abaxially with sparse red-brown glandular dots, orbicular or broadly elliptic. (34)	V. subalpinum
10 (8) +	Leaves abaxially without red-brown glandular dots, broadly elliptic to linear	(11)
11 (10)	Both surfaces of leaf blade glabrous	(12)
11 (10) +	Leaf blade abaxially stellate-pubescent in axils of veins at least, sometimes also along veins, adaxially glabrous or sparsely pubescent	(14)
12 (11)	Leaf margin remotely crenulate except at base, apex acute; corolla tube ca. 3.5 mm. (32)	V. omeiense
12 (11) +	Leaf margin remotely and shallowly serrate above base, apex abruptly narrowed or acuminate to long acuminate, shortly or long caudate; corolla tube 6-8 mm	(13)
13 (12)	Leaves subleathery to leathery, rarely thickly papery, midvein adaxially obviously raised, petiole purple-red; inflorescence peduncle (1.2-)2.5-7 cm; anthers reddish purple; fruit 6-7 mm. (31)	V. oliganthum
13 (12) +	Leaves papery, midvein adaxially not obviously raised, petiole green; inflorescence peduncle (3.5-)6-9 cm; anthers yellowish white; fruit 8-10 mm. (29)	V. longipedunculatum
14 (11)	Petiole glabrous, 2.5-5 cm; leaf blade 14-20 cm. (37)	V. trabeculosum
14 (11) +	Petiole stellate-pubescent, sparsely stellate-pubescent, or glabrous, less than 2.5 cm; leaf blade 2-14 cm	(15)
15 (14)	Petiole less than 10 mm	(16)
15 (14) +	Petiole 10-25 mm	(17)
16 (15)	Inflorescence 4-5 × 3-4 cm, rays sparsely stellate-pubescent, immediately glabrescent; anthers brownish purple; plant deciduous. (22)	V. brevitubum
16 (15) +	Inflorescence ca. 3 × 2 cm, rays stellate-pubescent; anthers yellowish white; plant evergreen. (35)	V. taitoense
17 (15)	Lateral veins mostly anastomosing near margin; leaves subleathery. (23)	V. chingii
17 (15) +	Lateral veins mostly directly ending in teeth; leaves papery	(18)
18 (17)	Corolla hypocrateriform, tube 5-6(-8) mm; stamens slightly exceeding corolla tube; flowers fragrant; leaf petiole purple. (25)	V. erubescens
18 (17) +	Corolla rotate-campanulate, tube ca. 3 mm; stamens obviously exceeding corolla tube; flowers not fragrant; leaf petiole green or reddish. (33)	V. shweliense
19 (1)	Winter buds naked; young branchlets and young leaves densely stellate-tomentose, fruit maturing from green, to red then black	(20)
19 (1) +	Winter buds with 1 or 2 (very rarely 3 or more) pairs of scales (rarely winter bud naked, and if such, buds, young branchlets, abaxial surfaces of leaves, inflorescences, calyces, corollas, and fruit with lepidote scales, see V. punctatum ); young branchlets and young leaves stellate-pubescent, bristlelike hairy, or glabrous, fruit maturing from green to red ( V. sect. Odontotinus and V. sect. Opulus ) or from green to blue-black ( V. sect. Tinus and V. sect. Megalotinus p.p.), rarely maturing from green, to red then black ( V. sect. Tomentosa and V. sect. Megalotinus p.p.)	(34)
20 (19)	Stipules 2 subulate, sometimes absent; inflorescence not pedunculate; pyrenes with 1 shallow dorsal groove and 1 deep ventral groove (2. V. sect. Pseudotinus )	(21)
20 (19) +	Stipules always absent; inflorescence pedunculate; pyrenes usually with 2 dorsal grooves and (1-)3 ventral grooves (1. V. sect. Viburnum )	(22)
21 (20)	Inflorescence without large sterile flowers, inflorescence rays and calyx tube with small brownish red glands; anthers purple; style exceeding calyx lobes. (14)	V. nervosum
21 (20) +	Inflorescence with large sterile flowers on margin, inflorescence rays and calyx tube without small brownish red glands; anthers yellow; style not exceeding calyx lobes. (15)	V. sympodiale
22 (20)	Shrubs or small trees, evergreen or semievergreen; leaf margin entire or indistinctly dentate, rarely serrulate; lateral veins often anastomosing near margin, not directly ending in teeth	(23)
22 (20) +	Leaves deciduous in winter, margin often dentate	(27)
23 (22)	Calyx tube glabrous; leaves 2-6(-8.5) cm, veins adaxially not impressed	(24)
23 (22) +	Calyx tube ± stellate-pubescent; leaves 5-25 cm	(25)
24 (23)	Corolla campanulate-funnelform, lobes shorter than tube; old leaves abaxially evenly stellate- pubescent, hairs not totally covering leaf surface; flowers fragrant; leaf lateral veins 3 or 4 pairs. (5)	V. congestum
24 (23) +	Corolla rotate, lobes as long as or slightly longer than tube; stellate hairs totally covering abaxial surfaces of old leaves; flowers not fragrant; leaf lateral veins 5 or 6 pairs. (13)	V. utile
25 (23)	Leaf blade adaxially densely stellate-pubescent, margin serrulate, apex acute to shortly acuminate, rarely obtuse to rounded. (1)	V. buddleifolium
25 (23) +	Leaf blade adaxially glabrous or glabrescent, sparsely stellate-pubescent when young particularly on midvein and lateral veins, margin entire or rarely inconspicuously dentate, apex slightly acute or obtuse	(26)
26 (25)	Leaves lanceolate-oblong to narrowly oblong, usually 5-15 × 1.5-4.5 cm, thickly papery when old; lateral veins and veinlets adaxially slightly impressed, not obviously rugose; petioles 1-2 cm; corolla outside sparsely stellate-pubescent; flowers on rays of 2nd order. (4)	V. chinshanense
26 (25) +	Leaves ovate-lanceolate to ovate-oblong, usually 8-25 × 2.5-8 cm, leathery; midvein, lateral veins, and veinlets adaxially deeply impressed, strongly rugose; petioles 1.5-4 cm; corolla outside subglabrous; flowers on rays of 3rd order. (10)	V. rhytidophyllum
27 (22)	Lateral veins directly ending in teeth, not anastomosing, or at least mostly so	(28)
27 (22) +	Lateral veins anastomosing near margin, not directly ending in teeth, or at least mostly so	(29)
28 (27)	Calyx tube glabrous; corolla lobes shorter than tube; stamens shorter than corolla; style exceeding calyx lobes; leaf margin serrulate or entire. (6)	V. cotinifolium
28 (27) +	Calyx tube stellate-tomentose; corolla lobes nearly as long as or less than 2 × as long as tube; stamens slightly exceeding corolla lobes; style not exceeding calyx lobes; leaf margin dentate. (7)	V. glomeratum
29 (27)	Inflorescence with large sterile radiant flowers. (8)	V. macrocephalum
29 (27) +	Inflorescence with only fertile flowers, without large sterile radiant flowers	(30)
30 (29)	Corolla rotate, tube shorter than lobes	(31)
30 (29) +	Corolla tubular-campanulate, rarely hypocrateriform, tube obviously longer than lobes	(32)
31 (30)	Branchlets of previous year brownish gray; leaves apically obtuse or rounded, sometimes emarginate or slightly acute; flowers mostly on rays of 3rd and 4th orders, flowers appearing with leaves; pyrenes dorsally raised or indistinctly 2-grooved. (11)	V. schensianum
31 (30) +	Branchlets of previous year yellow-white; leaves apically usually acute, rarely slightly obtuse; flowers mostly on rays of 2nd order, flowers appearing after leaves; pyrenes with 2 dorsal grooves. (2)	V. burejaeticum
32 (30)	Corolla hypocrateriform. (3)	V. carlesii
32 (30) +	Corolla tubular-campanulate	(33)
33 (32)	Leaves broadly ovate or elliptic to rarely suborbicular, 2.5-5(-6) cm, apex acute or obtuse; petioles 4-10 mm; flowers usually on rays of 1st order; corolla yellow-white. (9)	V. mongolicum
33 (32) +	Leaves ovate-lanceolate or ovate-oblong, 7-15(-18) cm, apex acuminate to narrowly caudate; petioles 1-4 cm; flowers on rays from 3rd to 4th order; corolla purple outside. (12)	V. urceolatum
34 (19)	Pyrenes globose, ovoid, or ellipsoid, with or without a shallow and narrow ventral groove, absolutely not compressed; fruit blue-black or turning from blue to black; leaves evergreen, glabrous or subglabrous (3. V. sect. Tinus )	(35)
34 (19) +	Pyrenes not as above; if ellipsoid then pyrenes with a deep ventral groove wide in upper part yet narrow in lower part; fruit red when mature, or turning from red to black, rarely yellow	(39)
35 (34)	Leaves pinnativeined. (16)	V. atrocyaneum
35 (34) +	Leaves triplinerved	(36)
36 (35)	Leaves apically obtuse to rounded, 2-6(-7.5) cm, margin entire; inflorescence peduncle ca. 1 cm. (20)	V. triplinerve
36 (35) +	Leaves apically acute to acuminate, margin entire or remotely serrate; inflorescence peduncle 1-3.5(-4) cm	(37)
37 (36)	Corolla white; old leaves thickly leathery; veinlets adaxially deeply impressed, obviously rugose; anthers reddish black. (18)	V. davidii
37 (36) +	Corolla greenish white or yellowish green; old leaves leathery; veinlets adaxially sometimes slightly impressed, not obviously rugose; anthers yellow	(38)
38 (37)	Shrubs 1-2 m tall; inflorescence 2-7 cm in diam., glabrous; leaves 3-9(-11) cm, margin usually remotely serrate. (19)	V. propinquum
38 (37) +	Shrubs or small trees 3-6 m tall; inflorescence 6-15 cm in diam.; leaves 6-13(-18) cm, margin entire or occasionally remotely serrate toward apex. (17)	V. cinnamomifolium
39 (34)	Winter buds surrounded by 2 pairs of connate scales; leaves 3-5-lobed; petiolar apex or leaf base adaxially with 1-4 conspicuous glands (8. V. sect. Opulus )	(40)
39 (34) +	Winter buds with 1 or 2 pairs of separate scales; glands absent at petiolar apex or leaf base adaxially, but abaxially sometimes with circular glands on both sides of midvein near base	(41)
40 (39)	Inflorescence without large sterile flowers; leaves 3-5-lobed; petioles not longer than 2.5 cm. (72)	V. koreanum
40 (39) +	Inflorescence with large sterile flowers on margin; leaves usually 3-lobed or those in upper part of branchlets undivided; petioles 1-5 cm. (73)	V. opulus
41 (39)	Leaves palmately 3-5-lobed, palmatinerved (7. V. sect. Odontotinus , p.p.). (62)	V. kansuense
41 (39) +	Leaves not divided or 2- or 3-lobed, mostly pinnativeined, sometimes 2 lateral veins at base appearing triplinerved	(42)
42 (41)	Inflorescence with large sterile flowers; fruit maturing from green to red then black; pyrenes with 1 ventral groove broad in upper part yet narrow in lower part (5. V. sect. Tomentosa )	(43)
42 (41) +	Inflorescence without large sterile flowers; fruit maturing from green to red, rarely to black (if so see V. cylindricum , V. lutescens , and V. punctatum ); pyrenes usually compressed, usually with shallow ventral and dorsal grooves	(44)
43 (42)	Leaves with more than 10 pairs of lateral veins; peduncle with 6-8 rays; inflorescences at apices of short lateral branches. (40)	V. plicatum
43 (42) +	Leaves with 5-7(-9) pairs of lateral veins; peduncle usually with 5 rays; inflorescence terminal. (39)	V. hanceanum
44 (42)	Shrubs or small trees evergreen, rarely deciduous (if so see V. ternatum ); stipules usually absent (but present in V. ternatum and V. inopinatum ); winter buds with a pair of separate scales, very rarely naked (if so see V. punctatum ) (6. V. sect. Megalotinus )	(45)
44 (42) +	Shrubs, rarely small trees, deciduous, rarely evergreen; stipules sometimes present; winter buds with 2 pairs of separate scales (7. V. sect. Odontotinus , p.p.)	(53)
45 (44)	Winter buds naked; winter buds, abaxial leaf surfaces, inflorescences, and outside of corollas with lepidote scales. (47)	V. punctatum
45 (44) +	Winter buds 2-perulate; plant not lepidote	(46)
46 (45)	Lateral veins totally or at least partly ending in teeth; inflorescence at apices of lateral short branchlets with 1-jugate leaves	(47)
46 (45) +	Lateral veins arched and anastomosing near margin, not directly ending in teeth; inflorescences at apices of branchlets with 1 to several pairs of leaves	(48)
47 (46)	Young branchlets, petioles, and inflorescences yellow-brown stellate-tomentose; first node of inflorescence with 6 or 7 rays. (41)	V. amplifolium
47 (46) +	Plant glabrous throughout; first node of inflorescence with 4 or 5 rays. (44)	V. laterale
48 (46)	Corolla campanulate; lobes short, erect; leaves abaxially with reddish or yellow glandular dots. (42)	V. cylindricum
48 (46) +	Corolla rotate	(49)
49 (48)	Leaf margin serrate, or sometimes serrate except at base, base not glandular spotted; stamens slightly exceeding corolla, filaments not folded in bud	(50)
49 (48) +	Leaf margins entire or sometimes slightly dentate at apex, base often glandular spotted on midvein; stamens obviously exceeding corolla, filaments folded in bud	(51)
50 (49)	Branchlets and abaxial leaf surfaces sparsely stellate-pubescent, glabrescent; plant elsewhere subglabrous; leaves leathery; inflorescence a compound umbel-like or panicle-like cyme, fragrant. (46)	V. lutescens
50 (49) +	Branchlets, abaxial leaf surfaces, petioles, peduncles, pedicels, and calyces yellow- brown stellate-tomentose; leaves papery; inflorescence a pyramidal panicle composed of 2-4 layers of umbels, not fragrant. (48)	V. pyramidatum
51 (49)	Leaves often whorled, 3 per whorl; stipules 2, persistent; inflorescence peduncle nearly absent. (49)	V. ternatum
51 (49) +	Leaves opposite; stipules absent or 2, caducous; inflorescence peduncle (1-)1.5-3 cm, rarely absent	(52)
52 (51)	Leaves pubescent or only so on veins, papery; calyx tube glabrous. (45)	V. leiocarpum
52 (51) +	Leaves thickly stellate-tomentose, leathery; calyx tube stellate-tomentose. (43)	V. inopinatum
53 (44)	Leaf blade oblong-lanceolate, lanceolate, or linear-lanceolate, 9-19(-27) × 1-4 cm, irregularly or remotely serrate, sometimes subentire; pyrenes compressed, subquadrangular or slightly rectangular	(54)
53 (44) +	Leaf blade orbicular, ovate to elliptic-oblong, obovate, or rhombic-ovate, rarely broadly oblong-lanceolate, generally less than 10 cm, margin dentate, serrate, denticulate, or entire, if more than 10 cm then more than 4 cm wide and with margin dentate, or less than 4 cm wide and with margin entire (see V. integrifolium ); pyrenes obovoid, ovoid, ellipsoid-ovoid, or ellipsoid	(55)
54 (53)	Young branchlets, leaves, petioles, and inflorescence axes glabrous; leaves with ca. 6 pairs of lateral veins. (71)	V. squamulosum
54 (53) +	Young branchlets, veins on abaxial leaf surfaces, petioles, and inflorescence axes stellate-pubescent; leaves with 7-12 pairs of lateral veins. (63)	V. lancifolium
55 (53)	Leaf margin entire, possibly remotely serrate or remotely dentate above middle part	(56)
55 (53) +	Leaf margin dentate, serrate, or denticulate, sometimes only so above 1/3 from base	(60)
56 (55)	Shrubs deciduous; inflorescence peduncle more than 2 cm	(57)
56 (55) +	Shrubs evergreen; inflorescence peduncle less than 2 cm or nearly absent	(58)
57 (56)	Leaves glabrous on both sides, margin entire, apex abruptly narrowed and long caudate, petiole green; style not exceeding calyx lobes. (61)	V. integrifolium
57 (56) +	Leaves stellate-pubescent at least abaxially, margin subentire, apex acute to shortly acuminate, petiole reddish purple; style exceeding calyx lobes. (56)	V. foetidum
58 (56)	Leaves abaxially with both golden yellow glandular dots and red-brown to black-brown glandular dots, adaxially usually not nigrescent when dry. (51)	V. chunii
58 (56) +	Leaves with black or brown glandular dots, nigrescent when dry	(59)
59 (58)	Calyx tube stellate-pubescent; leaves subleathery; fruit acute at apex. (59)	V. hainanense
59 (58) +	Calyx tube glabrous; leaves leathery; fruit rounded at apex. (69)	V. sempervirens
60 (55)	Leaves with 2-4 pairs of lateral veins, basal pair often appearing triplinerved	(61)
60 (55) +	Leaves with more than 5 pairs of pinnate lateral veins	(62)
61 (60)	Leaf blade 0.8-3 cm, apex rounded or slightly acute; inflorescence ca. 2.5 cm in diam., peduncles ca. 5 mm; pyrenes with 1 ventral groove. (68)	V. parvifolium
61 (60) +	Leaf blade 4-10 cm, apex acute to shortly acuminate; inflorescence 5-8 cm in diam., peduncles 2-5 cm; pyrenes with 3 ventral grooves. (56)	V. foetidum
62 (60)	Corolla outside glabrous or subglabrous, very rarely hairy in bud but later glabrescent (cf. V. betulifolium )	(63)
62 (60) +	Corolla sparsely or densely stellate-pubescent	(68)
63 (62)	Inflorescence or infructescence nodding; leaves nigrescent or slightly pale black when dry. (70)	V. setigerum
63 (62) +	Inflorescence or infructescence not nodding; leaves not nigrescent when dry	(64)
64 (63)	Inflorescence peduncles (5-)6-10(-12.5) cm. (60)	V. hengshanicum
64 (63) +	Inflorescence peduncles shorter than 5 cm	(65)
65 (64)	Petioles more than 1 cm; inflorescence 4-12 cm in diam	(66)
65 (64) +	Petiole less than 15 mm, inflorescence less than 4 cm in diam	(67)
66 (65)	Stipules caducous, or absent; first node of inflorescence usually with 5 rays; flowers on rays of 2nd and 3rd orders; fruit dark purple-red, maturing black, ellipsoid; branchlet of current year grayish black. (66)	V. melanocarpum
66 (65) +	Stipules persistent, often present; first node of inflorescence usually with 7 rays; flowers on rays from 3rd to 5th orders; fruit red, globose; branchlet of current year purple-brown. (50)	V. betulifolium
67 (65)	Stipules absent, petiole 5-15 mm, leaf blade ovate, apex caudate. (58)	V. formosanum
67 (65) +	Stipules present or absent, petiole 3-5 mm, leaf blade ovate-lanceolate, ovate- oblong, narrowly ovate, elliptic, or oblong-lanceolate, apex acuminate or acute. (55)	V. erosum
68 (62)	Leaves abaxially with yellow or yellowish or nearly colorless transparent glandular dots	(69)
68 (62) +	Leaves abaxially without glandular dots	(70)
69 (68)	Petiole 1-3 cm, stellate pubescent or simple hairs less than 1 mm; calyx tube stellate-pubescent, flowers fragrant; plant 2.5-5 m tall. (54)	V. dilatatum
69 (68) +	Petiole 0.5-1 cm, densely yellowish brown bristlelike hairy; calyx tube with simple hairs, flowers not fragrant; plant 1-2 m tall. (52)	V. corylifolium
70 (68)	Leaves adaxially with transparent or dispersed red-brown glandular dots	(71)
70 (68) +	Leaves adaxially without glandular dots	(72)
71 (70)	Leaf blade abaxially densely stellate-pubescent, adaxially with dispersed red- brown glandular dots; inflorescence peduncle 1-3.5 cm or very rarely nearly absent; pyrenes compressed, ovoid, ca. 6 × 4 mm, with 1 dorsal groove and 2 ventral grooves. (57)	V. fordiae
71 (70) +	Leaf blade abaxially sparsely stellate-pubescent or forklike pubescent, adaxially with transparent glandular dots; inflorescence peduncle usually very short or nearly absent, very rarely to 1.5 cm; pyrenes very compressed, ovoid, 4-5 × 3-4 mm, with 3 shallow dorsal grooves and 2 shallow ventral grooves. (65)	V. luzonicum
72 (70)	Leaf blade abaxially glabrous except midvein and lateral veins yellow-brown bristlelike hairy; lateral veins 8-12-jugate. (53)	V. dalzielii
72 (70) +	Leaf blade abaxially stellate-pubescent, or only so in vein axils; lateral veins 5-9-jugate	(73)
73 (72)	Styles very short, shorter than calyx lobes; corolla lobes nearly as long as tube; stamens shorter than corolla; pyrenes 4-6 mm; leaf margin serrate. (67)	V. mullaha
73 (72) +	Styles exceeding calyx lobes; corolla lobes longer than tube; stamens equaling or slightly exceeding corolla; pyrenes 6-7.5 mm; leaf margin crenate. (64)	V. longiradiatum

形态特征

灌木或乔木，有时带蔓性，落叶或常绿，常被簇状毛；冬芽裸露或有芽鳞。单叶，对生或稀3枚轮生；托叶有或无，或在同一个体上兼可存在或缺失。花序为由小聚伞花序集合而成的复伞形、圆锥形、有时近似伞房状圆锥形的混合花序，很少紧缩成簇状，有时周围或全部具白色大型的不孕花；花辐射对称，萼齿5，花冠辐状、钟状、漏斗状或高脚碟状，5裂；雄蕊5；子房1室，花柱极短，柱头浅3裂；胚珠1颗，自子房顶端下垂。果实为浆果状核果，顶端具宿存的萼齿及花柱；核多扁平，少有球形、卵状球形或椭圆状球形，内含1 颗种子；胚直，胚乳坚实或嚼烂状。约200种，分布于北半球温带和亚热带地区，尤其是亚洲及南美洲。我国约有80种，广布于各省区，尤以西南部种类最多；云南有43种，1亚种，10变种和1变型。本属植物有不少可入药及提取纤维用，少数种类之果实可供食用及酿酒，种子可榨油，个别的还是香料或蜜源植物。有些种类的花序具有白色而繁茂的小花和大型的不孕花以及鲜红夺目的果实，适于作园林绿化树种。

2. 荚蒾属——Viburnum Linn.

Linn. Sp. Pl. 267. 1753; Rehd. in Sarg. Trees and Shrubs 2: 105. 1908.

灌木或小乔木，落叶或常绿，常被簇状毛，茎干有皮孔。冬芽裸露或有鳞片。单叶，对生，稀3枚轮生，全缘或有锯齿或牙齿，有时掌状分裂，有柄；托叶通常微小，或不存在。花小，两性，整齐；花序由聚伞合成顶生或侧生的伞形式、圆锥式或伞房式，很少紧缩成簇状，有时具白色大型的不孕边花或全部由大型不孕花组成；苞片和小苞片通常微小而早落；萼齿5，宿存；花冠白色，较少淡红色，辐状、钟状、漏斗状或高脚碟状，裂片5枚，通常开展，很少直立，蕾时覆瓦状排列；雄蕊5枚，着生于花冠筒内，与花冠裂片互生，花药内向，宽椭圆形或近圆形；子房1室，花柱粗短，柱头头状或浅 (2-) 3裂；胚珠1颗，自子房顶端下垂。果实为核果，卵圆形或圆形，冠以宿存的萼齿和花柱；核扁平，较少圆形，骨质，有背、腹沟或无沟，内含1颗种子；胚直，胚乳坚实，硬肉质或嚼烂状。

全世界约有200种，分布于温带和亚热带地区；亚洲和南美洲种类较多。我国约有74种，广泛分布于全国各省区，以西南部种类最多。

新选模式种；绵毛荚蒾 V. lantana Linn.

分布区域

物种2000中文版
标本分布记录

标本份数

地图

卫星

矢量图

国家省级市级县级

区系物种

亚地区地区亚区分布区

分布物种

Sample Name:	Almont	Formation:	Sentinel Butte
Age:		Epoch (to):	Paleocene~
Stage:	Thanetian~	Contributer:
Ecology:		Climate:

locality name:	Almont	Province/State:	North Dakota
Continents:		county:	Morton
country:	United States	longitude-decimal degrees:	-101.445
latitude-decimal degrees:	46.912	elevation [m]:
Notes:	ETE Locality 180, Almont

Almont-Crane et al., 1990
title:	A Preliminary Survey of Fossil Leaves and Well-Preserved Reproductive Structures from the Sentinel Butte Formation (Paleocene) near Almont, North Dakota.
Year:	1990	Type:	book
Volume:	20	Issue:
Pages:		Publisher:
Location of publisher:		General:
Authors:	P. R. Crane, S.R. Manchester, and D.L. Dilcher

Sample Name:	Antelope Creek	Formation:	Golden Valley
Age:		Epoch (to):	Eocene~
Stage:	Ypresian~	Contributer:
Ecology:		Climate:

locality name:	Antelope Creek	Province/State:	North Dakota
Continents:	North America	county:	Stark
country:	United States	longitude-decimal degrees:	-102.75
latitude-decimal degrees:	46.75	elevation [m]:
Notes:	USNM 14068

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

Taxa

References an Palaeoclimate reconstruction

地质年代表

Sample Infomation

Locality

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References an Palaeoclimate reconstruction

地质年代表

Berga bei Sangershausen-Mai and Walther, 1988
title:	Die pliozaenen Floren von Thueringen, Deutsche Demokratische Republik
Year:	1988	Type:	journal
Volume:	7	Issue:
Pages:	55-297	Publisher:
Location of publisher:		General:
Authors:	D. H. Mai and H. Walther

Comox Coalfield-Bell, 1957
title:	Flora of the Upper Cretaceous Nanaimo Group of Vancouver Island, British Columbia
Year:	1957	Type:	journal
Volume:	293	Issue:
Pages:	1-84	Publisher:
Location of publisher:		General:
Authors:	W. A. Bell

Comox Coalfield-Bell, 1957
title:	Flora of the Upper Cretaceous Nanaimo Group of Vancouver Island, British Columbia
Year:	1957	Type:	journal
Volume:	293	Issue:
Pages:	1-84	Publisher:
Location of publisher:		General:
Authors:	W. A. Bell