学名(拉丁名) | Pimpinella |
中文名 | 茴芹属 |
命名人 | L. |
中文科名 | 伞形科 |
类型 | genus |
拉丁科名 | Umbelliferae |
学名(拉丁名) | Pimpinella |
中文名 | 茴芹属 |
中文科名 | 伞形科 |
类型 | genus |
拉丁科名 | Umbelliferae |
学名(拉丁名) | Pimpinella |
类型 | genus |
拉丁科名 | Apiaceae |
学名(拉丁名) | Pimpinella |
中文名 | 茴芹属 |
类型 | genus |
拉丁科名 | Apiaceae |
学名(拉丁名) | Pimpinella |
中文名 | 茴芹属 |
中文科名 | 伞形科 |
类型 | genus |
拉丁科名 | Apiaceae |
学名(拉丁名) | Pimpinella |
发表年份 | 1753(年) |
类型 | genus |
拉丁科名 | Apiaceae |
1 | 果实有毛;无萼齿。 (2) | 毛果组 Sect. Tragium |
1 | 果实无毛;萼齿明显或无。 (21) | 光果组 Sect. Tragoselinum |
2 | 一年生草本;无总苞片和小总苞片。 | 微毛茴芹 Pimpinella puberula |
2 | 多年生草本;通常有总苞片和小总苞片。 (3) | |
3 | 茎上部叶与基生叶同形,三出分裂或羽状分裂,裂片3-5;仅顶生花序有全育花,侧生花序多为不育花。 | 巍山茴芹 Pimpinella weishanensis |
3 | 茎上部叶和基生叶异形,3裂或1-2回羽状分裂;顶生花序和侧生花序的小花均能育。 (4) | |
4 | 花柱基圆锥形或短圆锥形。 (5) | |
4 | 花柱基近于垫状。 (19) | |
5 | 基生叶不分裂或三出分裂,稀羽状分裂。 (6) | |
5 | 基生叶1-2回羽状分裂或三出式2回羽状分裂。 (17) | |
6 | 一般为须根。 (7) | |
6 | 根长圆锥形或圆柱形。 (10) | |
7 | 植株被淡黄色柔毛,基生叶及茎中、下部叶不分裂;花瓣深紫色,基部有爪,背面无毛。 | 深紫茴芹 Pimpinella atropurpurea |
7 | 植株被微柔毛,基生叶及茎中、下部叶三出分裂或羽状分裂,稀不分裂;花瓣白色,基部楔形,背面有毛(异叶茴芹 P. diversifolia DC.) (8) | 异叶茴芹 Pimpinella diversifolia |
8 | 花瓣披针形,顶端全缘微弯曲。 | 尖瓣异叶茴芹 Pimpinella diversifolia var. angustipetala |
8 | 花瓣倒卵形,顶端凹缺,有内折小舌片。 (9) | |
9 | 无匍匐茎。 | 异叶茴芹(原变种) Pimpinella diversifolia var. diversifolia |
9 | 匍匐茎长3-20厘米。 | 走茎异叶茴芹 Pimpinella diversifolia var. stolonifera |
10 | 茎中、下部叶与基生叶异形(丽江茴芹除外)。 (11) | |
10 | 茎中、下部叶与基生叶同形。 (15) | |
11 | 基生叶三出分裂或羽状分裂;小总苞片等于或短于花柄。 | 重波茴芹 Pimpinella bisinuata |
11 | 基生叶通常不分裂;小总苞片等于或长于花柄。 (12) | |
12 | 伞辐密被柔毛;花瓣微带紫色。 | 丽江茴芹 Pimpinella rockii |
12 | 伞辐被疏柔毛或稍粗糙;花瓣白色。 (13) | |
13 | 基生叶肾形;花柄近等长。 | 肾叶茴芹 Pimpinella renifolia |
13 | 基生叶心形、心状披针形或近于长三角形;花柄不等长。 (14) | |
14 | 基生叶心形。 | 杏叶茴芹 Pimpinella candolleana |
14 | 基生叶心状披针形,或近于长三角形。 | 云南茴芹 Pimpinella yunnanensis |
15 | 伞辐4-10,长0.5-2.5厘米;小伞形花序有花6-10。 | 中甸茴芹 Pimpinella chungdienensis |
15 | 伞辐8-20,长1-4厘米;小伞形花序有花10-25。 (16) | |
16 | 基生叶不分裂;小总苞片短于花柄。 | 革叶茴芹 Pimpinella coriacea |
16 | 基生叶三出分裂;小总苞片等于或长于花柄。 | 藏茴芹 Pimpinella tibetanica |
17 | 伞辐极不等长,果期伞辐长可达7厘米或近无。 | 直立茴芹 Pimpinella smithii |
17 | 伞辐近等长,果期伞辐长2-3厘米。 (18) | |
18 | 基生叶1-2回羽状分裂;花柱基圆锥形,花柱长为花柱基的2-3倍。 | 城口茴芹 Pimpinella fargesii |
18 | 基生叶1-2回三出分裂;花柱基短圆锥形,花柱稍长于花柱基。 | 瘤果茴芹 Pimpinella tonkinensis |
19 | 分枝少,花序少数;无总苞片或偶有1片,小总苞片长于花柄。 | 木里茴芹 Pimpinella silvatica |
19 | 茎上部有多数纤细分枝,花序多数;无总苞片和小总苞片。 (20) | |
20 | 茎中、下部叶三出分裂,裂片卵形。 | 灰叶茴芹 Pimpinella grisea |
20 | 茎中、下部叶2回羽状分裂,裂片卵状披针形。 | 下曲茴芹 Pimpinella refracta |
21 | 无萼齿。 (22) | |
21 | 萼齿明显。 (34) | |
22 | 基生叶和茎下部叶不分裂。 | 少花茴芹 Pimpinella rubescens |
22 | 基生叶和茎下部叶1-2回三出分裂、三出式2-3回羽状分裂,或1-2回羽状分裂。 (23) | |
23 | 基生叶和茎下部叶三出分裂或羽状分裂。 (24) | |
23 | 基生叶和茎下部叶1-2回三出分裂,或三出式2-3回羽状分裂。 (29) | |
24 | 伞辐较少,2-6;小伞形花序有花2-8。 (25) | |
24 | 伞辐5-20;小伞形花序有花5-25。 (26) | |
25 | 无总苞片和小总苞片,果柄较短,长0.5-1厘米。 | 细软茴芹 Pimpinella flaccida |
25 | 偶有1总苞片,小总苞片2-3;果柄长1-1.5厘米。 | 锯边茴芹 Pimpinella serra |
26 | 伞辐10-20;无总苞片和小总苞片;果实长卵形。 (27) | |
26 | 伞辐5-10;有总苞片和小总苞片;果实卵形或卵球形。 (28) | |
27 | 基生叶2回羽状分裂,裂片线形,宽1-2毫米。 | 蛇床茴芹 Pimpinella cnidioides |
27 | 基生叶羽状分裂,裂片卵状披针形或卵形,宽5-20 (-40) 毫米。 | 羊红膻 Pimpinella thellungiana |
28 | 每两对小羽片之间相距1-3厘米,小羽片2-3裂;伞辐长达4厘米;花柱基近于垫状。 | 落新妇茴芹 Pimpinella astilbifolia |
28 | 每两对小羽片之间相距0.5-1.5厘米,小羽片不分裂;伞辐较短,长约2厘米;花柱基短圆锥形。 | 台湾茴芹 Pimpinella niitakayamensis |
29 | 须根;基生叶及茎下部叶三出式2-3回羽状分裂。 | 沼生茴芹 Pimpinella helosciadia |
29 | 根长圆锥形或圆柱形,基生叶及茎下部叶2回三出分裂,或三出式2回羽状分裂。 (30) | |
30 | 果柄细长,达10-12毫米。 | 尖叶茴芹 Pimpinella acuminata |
30 | 果柄较短,长约2-5毫米。 (31) | |
31 | 伞辐长2-3厘米;花瓣卵形或阔卵形;花柱基短圆锥形;花柱短。 (32) | |
31 | 伞辐长达6.5厘米,花瓣长圆形;花柱基圆锥形;花柱较长,约为果实长的1/2,或与果实近等长。 (33) | |
32 | 伞辐10-15;小伞形花序有花10-15;花瓣基部楔形或有短爪,顶端全缘微内弯。 | 尾尖茴芹 Pimpinella caudata |
32 | 伞辐4-8;小伞形花序有花5-10,花瓣基部楔形,顶端凹缺,有内折小舌片。 | 短柱茴芹 Pimpinella brachystyla |
33 | 无小总苞片;花瓣顶端全缘。 | 川鄂茴芹 Pimpinella henryi |
33 | 小总苞片2-5,线形;花瓣顶端全缘或微凹。 | 菱叶茴芹 Pimpinella rhomboidea |
34 | 基生叶和茎下部叶1-2回三出分裂,或近于1-2回羽状分裂。 (35) | |
34 | 基生叶和茎下部叶三出式2-4回羽状分裂,或2回三出分裂。 (39) | |
35 | 花瓣紫红色,基部有爪;花柱基圆锥形,或近于垫状;胚乳腹面微凹。 | 紫瓣茴芹 Pimpinella purpurea |
35 | 花瓣白色,基部楔形;花柱基圆锥形;胚乳腹面平直。 (36) | |
36 | 须根;基生叶和茎下部叶三出分裂,稀2回三出分裂,叶裂片卵形、阔卵形。 | 短果茴芹 Pimpinella brachycarpa |
36 | 根长圆锥形或支根须根状;基生叶和茎下部叶三出分裂或2回三出分裂,叶裂片卵形、长卵形或菱形。 (37) | |
37 | 根长圆锥形;基生叶的叶裂片卵形或菱形,花瓣顶端全缘,不凹陷。 | 景东茴芹 Pimpinella liiana |
37 | 支根须根状;茎生叶的叶裂片卵形或长卵形;花瓣顶端凹陷,有内折的小舌片。 (38) | |
38 | 通常无总苞片;果实卵形。 | 朝鲜茴芹 Pimpinella koreana |
38 | 总苞片3-5;果实长卵形。 | 具萼茴芹 Pimpinella calycina |
39 | 基生叶的末回裂片线形,长0.5-2.5厘米,宽0.1-0.2厘米。 | 谷生茴芹 Pimpinella valleculosa |
39 | 基生叶的末回裂片卵形或卵状披针形,长2-8厘米,宽1-4厘米。 (40) | |
40 | 基生叶和茎下部叶的末回裂片顶端尖尾状;伞辐长可达7厘米;果实较大,长约4毫米。 | 锐叶茴芹 Pimpinella arguta |
40 | 基生叶和茎下部叶的末回裂片顶端不呈尾状;伞辐较短,长2-3.5厘米;果实较小,长约2毫米。 | 辽冀茴芹 Pimpinella komarovi |
1 | Calyx teeth obsolete; fruit usually ornamented, hairy, papillose or granular, occasionally smooth. | (2) |
1 + | Calyx teeth conspicuous or obsolete; fruit glabrous. | (18) |
2 (1) | Annuals or biennials; bracteoles absent, rarely 1–4. | (3) |
2 (1) + | Perennials; bracteoles present. | (5) |
3 (2) | Stylopodium conic; fruit oblong-ovoid. (1) | Pimpinella anisum |
3 (2) + | Stylopodium low-conic; fruit ovoid-globose or cordate-ovoid. | (4) |
4 (3) | Rays 15–20, 15–40 mm; bracteoles absent (Xinjiang). (2) | Pimpinella puberula |
4 (3) + | Rays 3–5, 5–10 mm; bracteoles 2–4 (Sichuan, Yunnan). (3) | Pimpinella silvatica |
5 (2) | Basal and cauline leaves homomorphic, ternate or 1-pinnate, pinnae 2 pairs; flowers fertile in terminal umbels, mainly sterile in lateral umbels. (4) | Pimpinella kingdon-wardii |
5 (2) + | Basal and cauline leaves heteromorphic, simple, ternate or 1–2-pinnate; flowers fertile in terminal and lateral umbels. | (6) |
6 (5) | Root fibrous. | (7) |
6 (5) + | Taproot cylindrical or fusiform. | (8) |
7 (6) | Petals dark purple, glabrous, base shortly clawed. (5) | Pimpinella atropurpurea |
7 (6) + | Petals white, hairy abaxially, base cuneate. (6) | Pimpinella diversifolia |
8 (6) | Basal and lower leaves 1–2-pinnate or ternate-1–2-pinnate, rarely ternate. | (9) |
8 (6) + | Basal and lower leaves simple, 1-pinnate or ternate. | (12) |
9 (8) | Rays and pedicels extremely unequal, rays 2–70 mm. (14) | Pimpinella smithii |
9 (8) + | Rays and pedicels subequal or slightly unequal, rays 5–25(–30) mm. | (10) |
10 (9) | Styles 2–4 × stylopodium; fruit sparsely pubescent to glabrous; basal leaves 1–2-pinnate. (15) | Pimpinella fargesii |
10 (9) + | Styles 1–1.5 × stylopodium; fruit densely or sparsely pubescent; basal leaves 1-pinnate, or ternate, 1–2-pinnate. | (11) |
11 (10) | Rays 4–10, slightly unequal; fruit densely pubescent, basal leaves 1-pinnate, rarely ternate. (16) | Pimpinella chungdienensis |
11 (10) + | Rays 8–16, subequal; fruit sparsely pubescent, basal leaves ternate-1–2-pinnate. (17) | Pimpinella tonkinensis |
12 (8) | Basal and lower leaves ternate, rarely simple. | (13) |
12 (8) + | Basal and lower leaves usually simple. | (14) |
13 (12) | Bracteoles equal to or longer than pedicels; vittae 1 in each furrow. (12) | Pimpinella tibetanica |
13 (12) + | Bracteoles equal to or shorter than pedicels; vittae 1–4 in each furrow. (13) | Pimpinella bisinuata |
14 (12) | Leaf blade reniform-rotund, cordate-lanceolate or long triangular, rarely cordate-ovate. | (15) |
14 (12) + | Leaf blade cordate-ovate. | (16) |
15 (14) | Styles 1–1.5 × stylopodium; basal and lower leaves cordate-lanceolate or long triangular (Sichuan, Yunnan). (10) | Pimpinella yunnanensis |
15 (14) + | Styles ca. 3 × stylopodium; basal and lower leaves reniform-rotund (Hubei). (11) | Pimpinella renifolia |
16 (14) | Rays 6–15, 1–3 cm; bracteoles equal to or longer than flowers, densely hirsute. (7) | Pimpinella rockii |
16 (14) + | Rays (6–)10–25, 1.5–4(–6) cm; bracteoles shorter than or longer than flowers, glabrous. | (17) |
17 (16) | Styles 1–1.5 × stylopodium; fruit papillose-rugose. (8) | Pimpinella coriacea |
17 (16) + | Styles 2–3 × stylopodium; fruit finely granulate. (9) | Pimpinella candolleana |
18 (1) | Calyx teeth conspicuous. | (19) |
18 (1) + | Calyx teeth obsolete, rarely minute. | (28) |
19 (18) | Basal and lower leaves ternate-2-pinnate, 1–3-ternate or 3–4-pinnate; rays glabrous or hirsute. | (20) |
19 (18) + | Basal and lower leaves 1–2-ternate or 1–2-pinnate; rays glabrous or pubescent. | (23) |
20 (19) | Fruit oblong-ovoid; ultimate leaf segments linear. (41) | Pimpinella valleculosa |
20 (19) + | Fruit ovoid or cordate-ovoid; ultimate leaf segments ovate-lanceolate, ovate, broad-ovate or suborbicular. | (21) |
21 (20) | Ultimate leaf segments broad-ovate or suborbicular, 1–2 × 1–1.5 cm; styles ca. 1 × stylopodium; vittae 1 in each furrow. (42) | Pimpinella xizangensis |
21 (20) + | Ultimate leaf segments ovate-lanceolate or rhombic, 2–8 × 1–4 cm; styles 2–3 × stylopodium; vittae 3 in each furrow. | (22) |
22 (21) | Rays 2–7 cm; fruit ca. 4 × 3 mm; leaflets acuminate or caudate at the apex. (43) | Pimpinella arguta |
22 (21) + | Rays 2–3.5 cm; fruit ca. 2 × 1.5–2 mm; leaflets acuminate at the apex. (44) | Pimpinella komarovii |
23 (19) | Petals purple, base shortly clawed. | (24) |
23 (19) + | Petals white, base cuneate; seed face plane. | (25) |
24 (23) | Plants 30–80 cm; seed face slightly concave (Yunnan). (35) | Pimpinella purpurea |
24 (23) + | Plants 10–30 cm; seed face flat (Xizang). (36) | Pimpinella pimpinellisimulacrum |
25 (23) | Root fusiform; petals without incurved apex (Yunnan). (37) | Pimpinella liana |
25 (23) + | Roots fibrous; petals with an incurved apex (NE China). | (26) |
26 (25) | Leaves ternate, leaflets ovate or broad-ovate. (38) | Pimpinella brachycarpa |
26 (25) + | Leaves 1–2-ternate, leaflets ovate, oblong-ovate, or rhombic. | (27) |
27 (26) | Bracts absent, occasionally 2–3; fruit cordate-ovoid, ca. 1 × 0.8 mm. (39) | Pimpinella koreana |
27 (26) + | Bracts persistent, 3–5; fruit oblong-ovoid, ca. 5 × 2 mm. (40) | Pimpinella calycina |
28 (18) | All leaves simple or ternate; umbels 0.5–1 cm across, mostly lateral. (18) | Pimpinella rubescens |
28 (18) + | Basal and lower leaves 1–2-ternate, ternate-2–3-pinnate or 1–3-pinnate; umbels 2–10 cm across, terminal and lateral. | (29) |
29 (28) | Basal and lower leaves 1–2-ternate or 1–2-pinnate. | (30) |
29 (28) + | Basal and lower leaves 1–3-ternate or ternate-2–3-pinnate. | (37) |
30 (29) | Rays 2–6; umbellules 2–8-flowered; basal leaves 1–2-ternate or 1-pinnate, pinnae 2 pairs. | (31) |
30 (29) + | Rays 5–25; umbellules 5–25-flowered; basal leaves 1–3-pinnate, pinnae 3–6 pairs. | (34) |
31 (30) | Bracteoles 2–3; pedicels 10–15 mm in fruit (Anhui). (19) | Pimpinella serra |
31 (30) + | Bracteoles 0; pedicels 1–10 mm in fruit (Guizhou, Sichuan, Xizang, Yunnan). | (32) |
32 (31) | Plants glabrous; umbels 3–5 cm across, very lax; fruit ca. 2 mm, usually only one mericarp matures. (20) | Pimpinella refracta |
32 (31) + | Plants sparsely puberulent or pubescent; umbels 1–3.5 cm across, compact or open; fruit ca. 1 mm, both mericarps mature. | (33) |
33 (32) | Plants little-branched above; rays 2–6; style ca. 1 × stylopodium. (21) | Pimpinella flaccida |
33 (32) + | Plants several- or much-branched above; rays 4–5; style ca. 2 × stylopodium. (22) | Pimpinella grisea |
34 (30) | Bracts and bracteoles present; fruit cordate-ovoid; basal leaves 1-pinnate (Taiwan). (23) | Pimpinella niitakayamensis |
34 (30) + | Bracts and bracteoles absent; fruit oblong-ovoid; basal leaves 1–3-pinnate-pinnatifid. | (35) |
35 (34) | Basal leaves 1-pinnate, pinnae incised or lacerate-pinnatifid, ultimate segments ovate-lanceolate or ovate. (24) | Pimpinella thellungiana |
35 (34) + | Basal leaves 2–3-pinnate, ultimate segments linear or lanceolate. | (36) |
36 (35) | Styles ca. 3 × stylopodium; seed face plane; basal leaves 2-pinnate (NE China). (25) | Pimpinella cnidioides |
36 (35) + | Styles ca. 1 × stylopodium; seed face slightly concave; basal leaves 2–3-pinnate (Xizang). (26) | Pimpinella filipedicellata |
37 (29) | Root fibrous; basal and lower leaves ternate-2–3-pinnate. (27) | Pimpinella helosciadoidea |
37 (29) + | Taproot cylindrical or fusiform; basal and lower leaves 2-ternate or ternate-1–3-pinnate. | (38) |
38 (37) | Styles ca. equal to stylopodium. | (39) |
38 (37) + | Styles 2–3 × stylopodium. | (40) |
39 (38) | Rays 4–6(–8), extremely unequal; pedicels 2–3 mm in fruit. (28) | Pimpinella brachystyla |
39 (38) + | Rays 4–10(–13), subequal; pedicels 10–20 mm in fruit. (29) | Pimpinella acuminata |
40 (38) | Rays 5–8, 1.5–2.5 cm; basal leaves ternate-2–3-pinnate. (30) | Pimpinella nyingchiensis |
40 (38) + | Rays (5–)10–25, 3–6(–8) cm; basal leaves 2–3-ternate. | (41) |
41 (40) | Petals obcordate, apex incurved; basal leaves 2-ternate. (31) | Pimpinella caudata |
41 (40) + | Petals oblong-ovate, apex not incurved; basal leaves 2–3-ternate. | (42) |
42 (41) | Rays 5–7, 4–5 cm; basal leaves 3-ternate. (32) | Pimpinella triternata |
42 (41) + | Rays 10–25, 0.5–4(–6.5) cm; basal leaves 2-ternate. | (43) |
43 (42) | Bracteoles 1–2, or 0; terminal leaflets oblong-ovate or oblong-rhombic. (33) | Pimpinella henryi |
43 (42) + | Bracteoles 2–5; terminal leaflets broad-ovate or rhombic. (34) | Pimpinella rhomboidea |
多年生草本,稀一年生或二年生。根圆锥形,少须根。茎直立,常有分枝。叶柄基部具中等大小的膜质叶鞘;基生叶与茎生叶同形或异形,叶片不分裂、三出分裂或三出式羽状分裂;茎上部叶逐渐变小,常各式分裂。复伞形花序顶生和侧生;总苞片和小总苞片无或1—6,常为线形,稀披针形;伞辐近等长或不等长;小伞形花序常有多数花,稀2—4朵;萼齿不明显或为三角状披针形;花瓣白色,稀粉红色或紫红色,卵形或阔倒卵形,顶端凹陷,有内折小舌片,稀全缘;花柱基圆锥形或短圆锥形,花柱短或长于花柱基,向两侧反曲。果实卵形或宽卵形,稀长卵形,基部心形,被毛或光滑,果棱线形或不显著;分生果横切面五角形或近圆形,棱槽内油管1—4,合生面油管2—6;胚乳腹面平直或微回。约150种,主要分布于欧、亚、非三洲温带,美洲有少数种类(北美太平洋沿岸1种,南美数种)。我国40余种;云南有28种2变种。
45.茴芹属——Pimpinella L.
L. Sp. Pl. ed. 1. 263. 1753. ——Pimpinella L. gr. Spuriopimpinellade Boiss. in Bull. Soc. Bot. France 53: 428. 1906. ——Spuriopimpinella (de Boiss.) Kitagawa in Journ. Jap. Bot. 17(10): 558. 1941.
一年生、二年生或多年生草本。须根或有长圆锥形的主根。茎通常直立,稀匍匐,一般有分枝。叶柄长于或短于叶片,或与叶片近等长,基部有叶鞘;叶片不分裂、三出分裂、三出式羽状分裂或羽状分裂,裂片卵形、心形、披针形或线形;茎生叶与基生叶异形或同形,茎生叶向上逐渐简化变小,茎上部叶通常无柄,只有叶鞘。复伞形花序顶生和侧生,有或无总苞片及小总苞片,线形,稀披针形,全缘,偶有3裂;伞辐近等长、不等长或极不等长;小伞形花序通常有多数花,罕为2-4朵;萼齿通常不明显,或呈三角形、披针形;花瓣卵形、阔卵形或倒卵形,白色,稀为淡红色或紫色,基部一般为楔形,罕有爪,顶端凹陷,有内折小舌片,或全缘,并不内折,背面有毛或光滑;花柱基圆锥形、短圆锥形,稀为垫状,花柱一般长于花柱基,向两侧弯曲,或与花柱基近等长。果实卵形、长卵形或卵球形,基部心形,两侧扁压,有毛或无毛,果棱线形或不明显;分生果横剖面五角形或近圆形;每棱槽内油管1-4,合生面油管2-6;胚乳腹面平直或微凹;心皮柄2裂至中部或基部。
约150种,产欧、亚、非三洲,少数分布至美洲。我国有39种,2变种。
本属模式种:虎耳草茴芹 Pimpinella saxifraga L.