| 学名(拉丁名) | Dryopteris |
| 中文名 | 鳞毛蕨属 |
| 命名人 | Adanson |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 中文名 | 鳞毛蕨属 |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 中文名 | 鳞毛蕨属 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 中文名 | 鳞毛蕨属 |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 中文名 | 鳞毛蕨属 |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 发表年份 | 1763(年) |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Dryopteris |
| 类型 | genus |
| 1 | 叶片一回奇数羽状，顶生羽片与侧生羽片同形。 | 奇羽亚属 Subgen. Pycnopteris |
| 1 | 叶片一至四回羽裂或羽状，顶生羽片羽裂渐尖，不与侧生羽片同形。 (2) | |
| 2 | 鳞片扁平，即基部不为泡囊状。 (3) | 平鳞亚属 Subgen. Dryopteris |
| 2 | 蕨株除具扁平鳞片外，叶柄，叶轴尤其是羽轴和小羽轴具泡状或基部扩大，先端毛发状鳞片。 (15) | 泡鳞亚属 Subgen. Erythrovariae |
| 3 | 小羽片基部对称。 (4) | |
| 3 | 小羽片基部两侧不对称。 (12) | |
| 4 | 叶片一回羽状，羽片全缘、羽状浅裂或深裂。 (5) | |
| 4 | 叶片二回至三回羽状或四回羽裂 (8) | |
| 5 | 5.羽片全缘或有浅缺刻或浅裂达羽片之半，侧脉单一，不分又，基部一对或上侧一条叶脉较短，只伸达半途，袍子囊群有盖，少有无盖。 | 毛柄鳞毛蕨组 Sect. Hirtipedes |
| 5 | 羽片羽状浅裂至深裂，近二回羽状（至少基部数对如此），裂片上的叶脉通常二叉，少有单一。 (6) | |
| 6 | 孢子囊群盖大，膜质或螺壳状，成熟时仍笼罩孢子囊群。 | 大果鳞毛蕨组 Sect. Pandae |
| 6 | 孢子囊群盖小，成熟时不笼罩孢子囊群或往往脱落。 (7) | |
| 7 | 叶轴、羽轴及叶片下面被纤维状鳞毛。 | 纤维鳞毛蕨组 Sect. Fibrillosae |
| 7 | 叶轴、羽轴及叶片下面被平直披针形鳞片。 | 平鳞鳞毛蕨组 Sect. Dryoptoris |
| 8 | 叶片二回羽裂或基部为三回羽裂。 (9) | |
| 8 | 叶片三回羽裂或多数为三回羽裂至四回羽裂。 (10) | |
| 9 | 小羽片或裂片边缘通常有长刺状锯齿，孢子囊群盖小而膜质，边缘撕裂或啮蚀状。 | 高山鳞毛蕨组 Sect. Caespitosae |
| 9 | 小羽片或裂片边缘不具长刺状齿，孢子囊群盖骨质全缘。 | 半育鳞毛蕨组 Sect. Pallidae |
| 10 | 叶片长圆披针形或宽三角形，羽片下侧小羽片不比上侧的长。 | 边生鳞毛蕨组 Sect. Marginatae |
| 10 | 叶片五角形，羽片下侧小羽片比上侧的为长。 (11) | |
| 11 | 羽片具短柄，末回裂片边缘长刺状齿。 | 长尾鳞毛蕨组 Sect. Lophodium |
| 11 | 羽片具长柄，末回裂片边缘不具长刺状齿。 | 柄羽鳞毛蕨组 Sect. Aemulae |
| 12 | 叶片阔披针形，叶片基部的羽片上下两侧对称。 (13) | |
| 12 | 叶片基部的羽片不对称，即基部下侧小羽片特别伸长。 (14) | |
| 13 | 叶片三回羽裂，裂片先端具刺尖。 | 西域鳞毛蕨组 Sect. Remotae Fraser-Jenkins |
| 13 | 叶片二至四回羽裂，裂片先端不具刺尖。 | 华丽鳞毛蕨组 Sect. Splendentes |
| 14 | 叶片三至四回羽状；叶柄基部的鳞片线状披针形，皱曲，金黄色。 | 华丽鳞毛蕨组 Sect. Splendentes |
| 14 | 叶片二至四回羽状；叶柄基部的鳞片披针形或卵状披针形，棕色。 | 稀羽鳞毛蕨组 Sect. Nephrocystis |
| 15 | 基部羽片下侧的基部下侧小羽片通常特别伸长。 | 变异鳞毛蕨组 Sect. Variae |
| 15 | 叶片基部羽片的基部下侧小羽片不伸长或往往较短。 (16) | |
| 16 | 叶柄基部的鳞片披针形或阔披针形，棕色或淡棕色，叶柄至叶轴鳞片密。 | 泡鳞鳞毛蕨组 Sect. Erythrovariae |
| 16 | 叶柄基部鳞片狭披针形或线状披针形，黑色或黑棕色，叶柄中部向上叶轴的鳞片稀疏。 | 黑鳞鳞毛蕨组 Sect. Indusiatae |
| 1 (1) | Lamina once imparipinnate, apical pinna similar to lower ones (1. D. subg. Pycnopteris ) | (1) |
| 1 (1) + | Lamina once pinnate to quadripinnate or quinquepinnatifid, pinnae near apex gradually reduced to a pinnatifid apex | (2) |
| 2 (1) | Fronds with non-glandular hairs; indusia orbicular or reniform, inferior or superior (4. D. subg. Nothoperanema ) | (3) |
| 2 (1) + | Fronds without hairs or with glandular or non-glandular hairs; indusia reniform, superior | (6) |
| 3 (2) | Indusia superior (above sori), flat, and reniform. (19) | D. sect. Nothoperanema |
| 3 (2) + | Indusia inferior (beneath sori), globose or semiglobose | (4) |
| 4 (3) | Pinnae and pinnules of every order with or without a large cordate or ovate-lanceolate and often persistent scale at base; indusia membranous, semiglobose, indehiscent; lower portion of sporangiate stalk with a few multicellular clavate septate paraphyses; scales at base of stipe with multicellular clavate appendages on margin. (20) | D. sect. Acrophorus |
| 4 (3) + | Pinnae and pinnules of every order without a large cordate scale at base; indusia leathery, globose, often splitting into 2 or 3 valves upon maturity; lower portion of sporangiate stalk with single-celled clavate septate paraphyses or without any paraphyses; scales at base of stipe with single-celled clavate appendages on margin or without any appendages | (5) |
| 5 (4) | Sori sessile; lower portion of sporangiate stalk with single-celled clavate septate paraphyses; scales at base of stipe with entire margins. (21) | D. sect. Diacalpe |
| 5 (4) + | Sori stalked; lower portion of sporangiate stalk without any paraphyses; scales at base of stipe with single-celled clavate appendages on margin. (22) | D. sect. Peranema |
| 6 (2) | Scales bullate (broad-based scales with ciliate apices) on stipe, rachis, and especially on costa and costule; if scales flat then grooves of pinna rachis and pinnule rachis closed near their bases and fronds with multicellular non-glandular hairs ( D. sect. Dryopsis ), or frond segments anadromously arranged ( D. sect. Acrorumohra ), or stipe and rachis scales stiff and narrowly lanceolate and filiform ( D. liboensis ) (3. D. subg. Erythrovariae ) | (7) |
| 6 (2) + | Scales flat, not bullate on stipe and rachis (2. D. subg. Dryopteris ) | (11) |
| 7 (6) | Frond segments anadromously arranged. (17) | D. sect. Acrorumohra |
| 7 (6) + | Frond segments catadromously arranged | (8) |
| 8 (7) | Grooves of pinna rachis and pinnule rachis closed near their bases and fronds with multicellular non-glandular hairs. (18) | D. sect. Dryopsis |
| 8 (7) + | Grooves of pinna rachis and pinnule rachis connected near their bases and fronds without hairs | (9) |
| 9 (8) | Lamina pentagonal-ovate, usually tripinnate; lowest basiscopic pinnule on lowest pinna markedly longer than those above; pinnule caudate and acutely pointed. (16) | D. sect. Variae |
| 9 (8) + | Lamina lanceolate or ovate-lanceolate, usually pinnate to bipinnate; lowest basiscopic pinnule on lowest pinna not markedly longer than those above; pinnule not caudate, rounded | (10) |
| 10 (9) | Upper stipe with many small scales; scales on base of stipe lanceolate, brown or light brown. (14) | D. sect. Erythrovariae |
| 10 (9) + | Upper stipe glabrous; scales on base of stipe linear-lanceolate, black or very dark brown. (15) | D. sect. Indusiatae |
| 11 (6) | Segments asymmetrical at base | (12) |
| 11 (6) + | Segments symmetrical at base | (15) |
| 12 (11) | Lamina broadly lanceolate, basal pinnae symmetrical | (13) |
| 12 (11) + | Lamina triangular, basal pinnae asymmetrical, basiscopic basal pinnule proliferated | (14) |
| 13 (12) | Lamina tripinnatifid, with acute tooth on apex of segment. (10) | D. sect. Remotae |
| 13 (12) + | Lamina bi- to quadripinnatifid, without acute tooth on apex of segment. (11) | D. sect. Splendentes |
| 14 (12) | Lamina tri- to quadripinnate, stipe base scales filiform, golden. (12) | D. sect. Purpurascentes |
| 14 (12) + | Lamina bi- to tripinnate, stipe base scales lanceolate or ovate-lanceolate, brown. (13) | D. sect. Nephrocystis |
| 15 (11) | Lamina once pinnate, pinnae entire to deeply pinnatifid | (16) |
| 15 (11) + | Lamina twice pinnate or more dissected | (19) |
| 16 (15) | Pinnae entire to lobed, veins simple, basal veins of a segment (or at least acroscopic one) terminating ?halfway to segment margin. (1) | D. sect. Hirtipedes |
| 16 (15) + | Pinnae pinnatifid or becoming nearly twice pinnate (at least several basal pairs), veins forked, rarely simple, terminating at segment margin | (17) |
| 17 (16) | Indusia membranous or strombuliform, ?thick, surrounding sporangia even when ripe. (2) | D. sect. Pandae |
| 17 (16) + | Indusia not surrounding sporangia when ripe, sometimes deciduous | (18) |
| 18 (17) | Scales filiform or linear on rachis, costa, and abaxial surface of lamina. (3) | D. sect. Fibrillosae |
| 18 (17) + | Scales lanceolate on rachis, costa, and abaxial surface of lamina. (4) | D. sect. Dryopteris |
| 19 (15) | Lamina bipinnatifid, pinnules serrate | (20) |
| 19 (15) + | Lamina tripinnatifid or mostly tripinnatisect to quadripinnatifid | (21) |
| 20 (19) | Margin of pinnule or segment with long, acute teeth; indusia margin lacerate or erose. (5) | D. sect. Caespitosae |
| 20 (19) + | Margin of pinnule or segment without long, acute teeth; indusia entire, cartilaginous. (6) | D. sect. Pallidae |
| 21 (19) | Lamina oblong-lanceolate or broadly triangular, pinnae usually symmetrical. (7) | D. sect. Marginatae |
| 21 (19) + | Lamina pentagonal, pinnae asymmetrical, basiscopic pinnule of pinnae longer | (22) |
| 22 (21) | Pinnae shortly stalked, ultimate segments with long, acute teeth. (8) | D. sect. Lophodium |
| 22 (21) + | Pinnae long stalked, ultimate segments without long, acute teeth. (9) | D. sect. Aemulae |
土生，中型植物。根状茎短粗，直立或斜升，少有横卧，被鳞片；鳞片狭披针形。叶簇生，多数一型，少数二型。叶柄基部密被鳞片，鳞片形态各样，卵形、卵状披针形至狭披针形，顶端渐尖，边缘全缘或有齿，棕色、栗色或几为黑色，筛孔狭长而不透明。叶片形态亦多样，披针形、倒披针形、椭圆形、三角状卵形至五角形，一至四回羽状。侧生羽片多对。小羽片为下先出，末回小羽片或裂片的基部上侧呈耳状突起，齿尖无芒。叶脉分离，小脉单一或分叉，不达叶边。叶质地草质至革质，叶面通常无毛，叶背及叶轴、羽轴常被相当多的狭披针形、钻形或泡状鳞片，或被纤维状鳞毛；羽轴、小羽轴及主脉下面圆形，上面有纵沟。孢子囊群圆形，背生或有的顶生于小脉上；有囊群盖或偶无盖；囊群盖圆肾形，膜质或纸质，以深缺刻着生，全缘或罕有睫毛，宿存或早落；孢子囊近球形，有长柄，环带由14-20个增厚细胞组成。孢子椭圆形，周壁具褶皱，常形成片状或瘤状突起。染色体基数x=41。本属约有230余种，广布于世界各地，以亚洲大陆（特别是中国、喜马拉雅地区、日本、朝鲜）为分布中心，达到了最大的发展。中国现知有127种，为中国蕨类植物中大属之一。云南有83种，占国产种类的2/3。本属可分为3个亚属：奇羽亚属 (Pycnopteris)、平鳞亚属 (Dryopteris)；泡鳞亚属 (Erythrovariae)。其中：奇羽亚属和泡鳞亚属主要分布于东亚热带、亚热带，而平鳞亚属则主要分布中国 一 喜马拉雅亚区的亚高山、高山带。本属不仅为鳞毛蕨科的主干，而且似为三叉蕨科的祖先类型，它和该科的某些具有分离叶脉的属（如肋毛蕨属 Ctenitis, 轴脉蕨属 Ctenitopsis）在形体上经常相同，但叶干后不变褐棕色，而为淡绿色，小羽轴上面不隆起，也不被棕色腊肠形的粗毛，而为凹陷成一纵沟，并且光滑无毛，囊群盖较大，往往为革质或角质，故易区别。Holttum (在 British Fern Gazette 9: part. 6. 208, 1965) 说“鳞毛蕨属与耳蕨属的分野在中国可能消失了”，这是没有根据的。这两个都是很自然的属，不存在中间类型。
7. 鳞毛蕨属* Dryopteris Adanson
Adanson, Fam. Des. Plantes 2: 20. 1763; Schott. Gen. Fil. ad t. 9. 1835: Gray, Manu. Bot. ed. 1. 631. 1848: Ching in Bull Fan Mem. Inst. Biol. Bot. 8: 363. 1938; H. Ito in Nakai et Honda, Nov. Fl. Jap. no 4. 1939: Cop. Gen. Fil. 121. 1947, excl. Diclisodon Moore et Microchlaena Ching; Fraser-Jenkins in Bull. Br. Mus. Nat. Hist. Bot. 14(3): 183-218. 1986 et 18(5): 323-477. 1989: S. G. Lu in ActaPhytotax. Sin. 31(4): 385-391. 1993. ——Psidopodium Necker, Elem. Bot. 3: 315. 1790 (fide Christensen. 1906: 590); ——Nephrodium Rich. in Marthe ex Michaux, Fl. Bor. Amer. 2: 266 (1803).
陆生中型蕨物。根状茎粗短，直立或斜升（偶为横走），顶端密被鳞片，鳞片卵形、阔披针形、卵状披针形或披针形，红棕色、褐棕色或黑色，有光泽，全缘、略有疏齿牙或呈流苏状，质厚，由狭长而不透明的细胞组成，胞壁厚而曲折。叶簇生，螺旋状排列、向四面放射呈中空的倒圆锥形，有柄，被同样的鳞片（有时叶柄基部以上几无鳞片）；叶片阔披针形、长圆形、三角状卵形、有时五角形，一回羽状或二至四回羽状或四回羽裂，顶部羽裂，罕为一回奇数羽状，如为多回羽状复叶，则分枝图式为除基部1对羽片的一回小羽片为上先出外，其余均为下先出，不被针状毛，但通常多少有鳞片（罕有光滑），鳞片披针形、线形、卷发形纤维状或基部为泡囊形，或基部扩大向顶端为钻状，全缘，或为流苏状，间为撕裂； 末回羽片基部圆形对称，罕为不对称的楔形（但基部上侧从不为耳状凸起），边缘通常有锯齿，少有具针状刺头。叶通常为纸质至近革质，少为草质，干后淡绿色或草绿色； 各回小羽轴或（或主脉）以锐角斜出，基部以狭翅下沿于下一回的小羽轴，下面圆形隆起，上面具纵沟，两侧具隆起的边，光滑无毛，且与下一回的小羽轴上面的纵沟互通。叶脉分离，羽状，单一或二至三叉，不达叶边，先端往往有明显的膨大水囊。孢子囊群圆形，生于叶脉背部，或罕有生于叶脉顶部、通常有囊群盖（少为无盖）；盖为圆肾形，通常大而全缘、光滑（偶有腺体或边缘啮蚀），棕色，平坦或有时为螺壳形，并笼罩整个孢子囊群，质较厚，有时为角质，宿存，以深缺刻着生于叶脉。孢子两面形，肾形或肾状椭圆形，表面有疣状突起或有阔翅状的周壁。染色体基数 n＝41。
模式种：欧洲鳞毛蕨 Polypodium filix-max L.（＝Dryopteris filix-mas (L.) Schott）
本属是自然的分类群，约有230余种，广布于世界各地，以亚洲大陆（特别是中国及喜马拉雅地区其他国家、日本、朝鲜）为分布中心，达到了最大的发展；中国现知有127种，为中国蕨类蕨物中大属之一。
本属不仅为鳞毛蕨科的主干，而且似为三叉蕨科的祖先类型，它和该科的某些具有分离叶脉的属（如肋毛蕨属 Ctenitis，轴脉蕨属 Ctenitopsis）在形体上经常相同，但叶干后不变褐棕色，而为淡绿色，小羽轴上面不隆起，也不被棕色腊肠形的粗毛，而为凹陷成一纵沟，并且光滑无毛，囊群盖较大，往往为革质或角质，故易区别。Holttum（在 British Fern Gazette 9: part. 6. 208. 1965）说“鳞毛蕨属与耳蕨属的分野在中国可能消失了”，这是没有根据的。这两个都是很自然的属，不存在中间类型。
本属可分为3个亚属：奇羽亚属（Subgen. Pycnopteris）、平鳞亚属（Subgen. Dry-opteris）；泡鳞亚属（Subgen. Erythrovariae）。
* 秦仁昌教授是中国蕨类植物学的奠基者，也是最早系统研究亚洲鳞毛蕨属的学者之一。早在1938年他就发表了《中国与印度及其邻邦产鳞毛蕨属（Dryopteris）之正误研究》（Bull. Fan. Mem. Inst. Biol. Bot. 8: 364-507. 1938），其中记载中国鳞毛蕨属植物87种11变种。60年代以来他又与裘佩熹、刘正宇、张朝芳等人在刊物和地方植物志中发表了近200个新种，并编制了本属的检索表。我们在编写本志时曾参考了他的手稿。限于当时的条件，他发表新种时所依据的标本甚少，往往只有一份。现据较多的标本，对他发表的新种有所归并。——编著者
| Sample Name: | Florissant, Denver Museum | Formation: | Florissant |
| Age: | Epoch (to): | Oligocene~Late/Upper Eocene | |
| Stage: | ~ | Contributer: | |
| Ecology: | Climate: | ||
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| locality name: | Florissant Flora | Province/State: | Colorado |
| Continents: | county: | ||
| country: | United States | longitude-decimal degrees: | -105.25 |
| latitude-decimal degrees: | 38.916667 | elevation [m]: | |
| Notes: | ETE Locality 1468 | ||
| Florissant Flora-MacGinitie, 1953 | |||
| title: | Fossil Plants of the Florissant Beds of Colorado | ||
| Year: | 1953 | Type: | journal |
| Volume: | 599 | Issue: | |
| Pages: | 1-198 | Publisher: | |
| Location of publisher: | General: | ||
| Authors: | H. D. MacGinitie | ||
Phanerozoic |
Proterozoic |
Archean |
Hadean (informal) |
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Cenozoic |
Mesozoic |
Paleozoic |
Neoproterozoic |
Mesoproterozoic |
Paleoproterozoic |
Neoarchean |
Mesoarchean |
Paleoarchean |
Eoarchean |
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Quaternary |
Neogene |
Paleogene |
Cretaceous |
Jurassic |
Triassic |
Permian |
Carboniferous |
Devonian |
Silurian |
Ordovician |
Cambrian |
Ediacaran |
Cryogenian |
Tonian |
Stenian |
Ectasian |
Calymmian |
Statherian |
Orosirian |
Rhyacian |
Siderian |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Holocene |
Pleistocene |
Pliocene |
Miocene |
Oligocene |
Eocene |
Paleocene |
Upper |
Lower |
Upper |
Middle |
Lower |
Upper |
Middle |
Lower |
Lopingian |
Guadalupian |
Cisuralian |
Pennsylvanian-Upper |
Pennsylvanian-Middle |
Pennsylvanian-Lower |
Mississippian-Upper |
Mississippian-Middle |
Mississippian-Lower |
Upper |
Middle |
Lower |
Pridoli |
Ludlow |
Wenlock |
Llandovery |
Upper |
Middle |
Lower |
Furongian |
Series 3 |
Series 2 |
Terreneuvian |
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Upper |
"lonia" |
Calabrian |
Gelasian |
Piacenzian |
Zanclean |
Messinian |
Tortonian |
Serravallian |
Langhian |
Burdigalian |
Chattian |
Aquitanian |
Rupelian |
Priabonian |
Bartonian |
Lutetian |
Ypresian |
Thanetian |
Selandian |
Danian |
Maastrichtian |
Campanian |
Santonian |
Coniacian |
Turonian |
Cenomanian |
Albian |
Aptian |
Barremian |
Hauterivian |
Valanginian |
Berriasian |
Tithonian |
Kimmeridgian |
Oxfordian |
Callovian |
Bathonian |
Bajocian |
Aalenian |
Toarcian |
Pliensbachian |
Sinemurian |
Hettangian |
Rhaetian |
Norian |
Carnian |
Ladinian |
Anisian |
Olenekian |
Induan |
Changhsingian |
Wuchiapingian |
Capitanian |
Wordian |
Roadian |
Kungurian |
Artinskian |
Sakmarian |
Asselian |
Gzhelian |
Kasimovian |
Moscovian |
Bashkirian |
Serpukhovian |
Visean |
Tournaisian |
Famennian |
Frasnian |
Givetian |
Eifelian |
Emsian |
Pragian |
Lochkovian |
Ludfordian |
Gorstian |
Homerian |
Sheinwoodian |
Telychian |
Aeronian |
Rhuddanian |
Hirnantian |
Katian |
Sandbian |
Darriwilian |
Dapingian |
Floian |
Tremadocian |
Stage 10 |
Stage 9 |
Paibian |
Guzhangian |
Drumian |
Stage 5 |
Stage 4 |
Stage 3 |
Stage 2 |
Fortunian |
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| 0 | 0.0117 | 0.126 | 0.781 | 1.806 | 2.588 | 3.6 | 5.332 | 7.246 | 11.608 | 13.82 | 15.97 | 20.43 | 23.03 | 28.4 ±0.1 | 33.9 ±0.1 | 37.2 ±0.1 | 40.4 ±0.2 | 48.6 ±0.2 | 55.8 ±0.2 | 58.7 ±0.2 | 61.1 | 65.5 ±0.3 | 70.6 ±0.6 | 83.5 ±0.7 | 85.8 ±0.7 | 88.6 | 93.6 ±0.8 | 99.6 ±0.9 | 112.0 ±1.0 | 125.0 ±1.0 | 130.0 ±1.5 | 133.9 | 140.2 ±3.0 | 145.5 ±4.0 | 150.8 ±4.0 | 155.6 | 161.2 ±4.0 | 164.7 ±4.0 | 167.7 ±3.5 | 171.6 ±3.0 | 175.6 ±2.0 | 183.0 ±1.5 | 189.6 ±1.5 | 196.5 ±1.0 | 199.6 ±0.6 | 203.6 ±1.5 | 216.5 ±2.0 | 228.7 | 237.0 ±2.0 | 245.9 | 249.5 | 251.0 ±0.4 | 253.8 ±0.7 | 260.4 ±0.7 | 265.8 ±0.7 | 268.0 ±0.7 | 270.6 ±0.7 | 275.6 ±0.7 | 284.4 ±0.7 | 294.6 ±0.8 | 299.0 ±0.8 | 303.4 ±0.9 | 307.2 ±1.0 | 311.7 ±1.1 | 318.1 ±1.3 | 328.3 ±1.6 | 345.3 ±2.1 | 359.2 ±2.5 | 374.5 ±2.6 | 385.3 ±2.6 | 391.8 ±2.7 | 397.5 ±2.7 | 407.0 ±2.8 | 411.2 ±2.8 | 416.0 ±2.8 | 418.7 ±2.7 | 421.3 ±2.6 | 422.9 ±2.5 | 426.2 ±2.4 | 428.2 ±2.3 | 436.0 ±1.9 | 439.0 ±1.8 | 443.7 ±1.5 | 445.6 ±1.5 | 455.8 ±1.6 | 460.9 ±1.6 | 468.1 ±1.6 | 471.8 ±1.6 | 478.6 ±1.7 | 488.3 ±1.7 | 492 | 496 | 499 | 503 | 506.5 | 510 | 515 | 521 | 528 | 542.0 ±1.0 | 635 | 850 | 1000 | 1200 | 1400 | 1600 | 1800 | 2050 | 2300 | 2500 | 2800 | 3200 | 3600 | 4000 | 4600 |
显生宇 |
元古宇 |
太古宇 |
冥古宇 (非正式) |
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新生界 |
中生界 |
古生界 |
新元古界 |
中元古界 |
古元古界 |
新太古界 |
中太古界 |
古太古界 |
始太古界 |
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第四系 |
新近系 |
古近系 |
白垩系 |
侏罗系 |
三叠系 |
三叠系 |
石炭系 |
泥盆系 |
志留系 |
奥陶系 |
寒武系 |
埃迪卡拉系 |
成冰系 |
拉伸系 |
狭带系 |
延展系 |
盖层系 |
固结系 |
造山系 |
层侵系 |
成铁系 |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
全新统 |
更新统 |
上新统 |
中新统 |
渐新统 |
始新统 |
古新统 |
上白垩统 |
下白垩统 |
上侏罗统 |
中侏罗统 |
下侏罗统 |
上三叠统 |
中三叠统 |
下三叠统 |
乐平统 |
瓜德鲁普统 |
乌拉尔统 |
宾夕法尼亚亚系-上 |
宾夕法尼亚亚系-中 |
宾夕法尼亚亚系-下 |
密西西比亚系-上 |
密西西比亚系-中 |
密西西比亚系-下 |
上泥盆统 |
中泥盆统 |
下泥盆统 |
普里道利统 |
罗德洛统 |
文洛克统 |
兰多维列统 |
上奥陶统 |
中奥陶统 |
下奥陶统 |
芙蓉统 |
第三统 |
第二统 |
纽芬兰统 |
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上更新统 |
“伊奥尼雅阶” |
卡拉布里雅阶 |
格拉斯阶 |
皮亚琴察阶 |
赞克尔阶 |
梅辛阶 |
托尔通阶 |
塞拉瓦尔阶 |
兰哥阶 |
布尔迪加尔阶 |
阿启坦阶 |
夏特阶 |
鲁培尔阶 |
普利亚本阶 |
巴尔通阶 |
鲁帝特阶 |
伊普里斯阶 |
坦尼特阶 |
赛兰特阶 |
丹尼阶 |
马斯特里赫特阶 |
坎潘阶 |
三冬阶 |
康尼亚克阶 |
土伦阶 |
赛诺曼阶 |
阿尔布阶 |
阿普特阶 |
巴雷姆阶 |
欧特里夫阶 |
凡兰吟阶 |
贝利阿斯阶 |
提塘阶 |
基末利阶 |
牛津阶 |
卡洛夫阶 |
巴通阶 |
巴柔阶 |
阿林阶 |
土阿辛阶 |
普林斯巴阶 |
辛涅缪尔阶 |
赫塘阶 |
瑞替阶 |
诺利阶 |
卡尼阶 |
拉丁阶 |
安尼阶 |
奥伦尼克阶 |
印度阶 |
长兴阶 |
吴家坪阶 |
卡匹敦阶 |
沃德阶 |
罗德阶 |
空谷阶 |
亚丁斯克阶 |
萨克马尔阶 |
阿瑟尔阶 |
格舍尔阶 |
卡西莫夫阶 |
莫斯科阶 |
巴什基尔阶 |
谢尔普霍夫阶 |
维宪阶 |
杜内阶 |
法门阶 |
弗拉阶 |
吉维特阶 |
艾菲尔阶 |
埃姆斯阶 |
布拉格阶 |
洛霍考夫阶 |
卢德福特阶 |
高斯特阶 |
侯墨阶 |
申伍德阶 |
特列奇阶 |
埃隆阶 |
鲁丹阶 |
赫南特阶 |
凯迪阶 |
桑比阶 |
达瑞威尔阶 |
大坪阶 |
弗洛阶 |
特马豆克阶 |
第十阶 |
第九阶 |
排碧阶 |
古丈阶 |
鼓山阶 |
第五阶 |
第四阶 |
第三阶 |
第二阶 |
幸运阶 |
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| 0 | 0.0117 | 0.126 | 0.781 | 1.806 | 2.588 | 3.6 | 5.332 | 7.246 | 11.608 | 13.82 | 15.97 | 20.43 | 23.03 | 28.4 ±0.1 | 33.9 ±0.1 | 37.2 ±0.1 | 40.4 ±0.2 | 48.6 ±0.2 | 55.8 ±0.2 | 58.7 ±0.2 | 61.1 | 65.5 ±0.3 | 70.6 ±0.6 | 83.5 ±0.7 | 85.8 ±0.7 | 88.6 | 93.6 ±0.8 | 99.6 ±0.9 | 112.0 ±1.0 | 125.0 ±1.0 | 130.0 ±1.5 | 133.9 | 140.2 ±3.0 | 145.5 ±4.0 | 150.8 ±4.0 | 155.6 | 161.2 ±4.0 | 164.7 ±4.0 | 167.7 ±3.5 | 171.6 ±3.0 | 175.6 ±2.0 | 183.0 ±1.5 | 189.6 ±1.5 | 196.5 ±1.0 | 199.6 ±0.6 | 203.6 ±1.5 | 216.5 ±2.0 | 228.7 | 237.0 ±2.0 | 245.9 | 249.5 | 251.0 ±0.4 | 253.8 ±0.7 | 260.4 ±0.7 | 265.8 ±0.7 | 268.0 ±0.7 | 270.6 ±0.7 | 275.6 ±0.7 | 284.4 ±0.7 | 294.6 ±0.8 | 299.0 ±0.8 | 303.4 ±0.9 | 307.2 ±1.0 | 311.7 ±1.1 | 318.1 ±1.3 | 328.3 ±1.6 | 345.3 ±2.1 | 359.2 ±2.5 | 374.5 ±2.6 | 385.3 ±2.6 | 391.8 ±2.7 | 397.5 ±2.7 | 407.0 ±2.8 | 411.2 ±2.8 | 416.0 ±2.8 | 418.7 ±2.7 | 421.3 ±2.6 | 422.9 ±2.5 | 426.2 ±2.4 | 428.2 ±2.3 | 436.0 ±1.9 | 439.0 ±1.8 | 443.7 ±1.5 | 445.6 ±1.5 | 455.8 ±1.6 | 460.9 ±1.6 | 468.1 ±1.6 | 471.8 ±1.6 | 478.6 ±1.7 | 488.3 ±1.7 | 492 | 496 | 499 | 503 | 506.5 | 510 | 515 | 521 | 528 | 542.0 ±1.0 | 635 | 850 | 1000 | 1200 | 1400 | 1600 | 1800 | 2050 | 2300 | 2500 | 2800 | 3200 | 3600 | 4000 | 4600 |
| Sample Name: | Florissant, U of California Loc. 3731 | Formation: | Florissant |
| Age: | Epoch (to): | Oligocene~Late/Upper Eocene | |
| Stage: | ~ | Contributer: | |
| Ecology: | Climate: | ||
|
|
|||
| locality name: | Florissant Flora | Province/State: | Colorado |
| Continents: | county: | ||
| country: | United States | longitude-decimal degrees: | -105.25 |
| latitude-decimal degrees: | 38.916667 | elevation [m]: | |
| Notes: | ETE Locality 1468 | ||
| Florissant Flora-MacGinitie, 1953 | |||
| title: | Fossil Plants of the Florissant Beds of Colorado | ||
| Year: | 1953 | Type: | journal |
| Volume: | 599 | Issue: | |
| Pages: | 1-198 | Publisher: | |
| Location of publisher: | General: | ||
| Authors: | H. D. MacGinitie | ||
Phanerozoic |
Proterozoic |
Archean |
Hadean (informal) |
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Cenozoic |
Mesozoic |
Paleozoic |
Neoproterozoic |
Mesoproterozoic |
Paleoproterozoic |
Neoarchean |
Mesoarchean |
Paleoarchean |
Eoarchean |
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Quaternary |
Neogene |
Paleogene |
Cretaceous |
Jurassic |
Triassic |
Permian |
Carboniferous |
Devonian |
Silurian |
Ordovician |
Cambrian |
Ediacaran |
Cryogenian |
Tonian |
Stenian |
Ectasian |
Calymmian |
Statherian |
Orosirian |
Rhyacian |
Siderian |
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Holocene |
Pleistocene |
Pliocene |
Miocene |
Oligocene |
Eocene |
Paleocene |
Upper |
Lower |
Upper |
Middle |
Lower |
Upper |
Middle |
Lower |
Lopingian |
Guadalupian |
Cisuralian |
Pennsylvanian-Upper |
Pennsylvanian-Middle |
Pennsylvanian-Lower |
Mississippian-Upper |
Mississippian-Middle |
Mississippian-Lower |
Upper |
Middle |
Lower |
Pridoli |
Ludlow |
Wenlock |
Llandovery |
Upper |
Middle |
Lower |
Furongian |
Series 3 |
Series 2 |
Terreneuvian |
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Upper |
"lonia" |
Calabrian |
Gelasian |
Piacenzian |
Zanclean |
Messinian |
Tortonian |
Serravallian |
Langhian |
Burdigalian |
Chattian |
Aquitanian |
Rupelian |
Priabonian |
Bartonian |
Lutetian |
Ypresian |
Thanetian |
Selandian |
Danian |
Maastrichtian |
Campanian |
Santonian |
Coniacian |
Turonian |
Cenomanian |
Albian |
Aptian |
Barremian |
Hauterivian |
Valanginian |
Berriasian |
Tithonian |
Kimmeridgian |
Oxfordian |
Callovian |
Bathonian |
Bajocian |
Aalenian | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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