学名(拉丁名) | Ajania |
中文名 | 亚菊属 |
命名人 | Poljak. |
中文科名 | 菊科 |
类型 | genus |
拉丁科名 | Compositae |
学名(拉丁名) | Ajania |
中文名 | 亚菊属 |
中文科名 | 菊科 |
类型 | genus |
拉丁科名 | Compositae |
学名(拉丁名) | Ajania |
中文名 | 亚菊属 |
类型 | genus |
拉丁科名 | Asteraceae |
学名(拉丁名) | Ajania |
中文名 | 亚菊属 |
中文科名 | 菊科 |
类型 | genus |
拉丁科名 | Asteraceae |
1 | 总苞麦秆黄色,有光泽,直径2-4毫米,总苞片边缘白色膜质。 (2) | |
1 | 总苞非麦秆黄色,无光泽,直径通常4-10毫米;总苞片边缘棕褐色、黑褐色、黑紫色或褐色。 (9) | |
2 | 头状花序5-10个在枝端排成规则的束状伞房花序;总苞圆柱状,总苞片硬草质,顶端急尖。 | 束伞亚菊 Ajania parviflora |
2 | 头状花序多数在茎枝顶端排成伞房或复伞房花序;总苞钟状,总苞片草质,顶端钝或圆。 (3) | |
3 | 小灌木或小半灌木。 (4) | |
3 | 多年生草本,有地下匍茎。 (8) | |
4 | 叶两面绿色或淡绿色,无毛或几无毛;末回裂片丝状。 | 丝裂亚菊 Ajania nematoloba |
4 | 叶两面或下面灰白色,被顺向贴伏的短柔毛;边缘锯齿或浅裂,末回裂片椭圆形、长椭圆形、线形、披针形、三角形或镰刀形。 (5) | |
5 | 叶边缘浅锯齿或浅裂。 | 川甘亚菊 Ajania potaninii |
5 | 叶二回羽状分裂或二回掌式或掌式羽状3-5全裂。 (6) | |
6 | 叶二回羽状分裂。 | 蓍状亚菊 Ajania achilloides |
6 | 叶二回掌式或掌式羽状3-5全裂。 (7) | |
7 | 全部叶的叶耳1-2回分裂,有长1-2毫米的叶耳柄。 | 光苞亚菊 Ajania nitida |
7 | 全部叶的叶耳1回分裂,无叶耳柄,因此叶耳贴茎或抱茎。 | 灌木亚菊 Ajania fruticulosa |
8 | 头状花序大,总苞宽6一7毫米,外层苞片长椭圆形,长2.5-3毫米,顶端有圆形蜡质附属物。 | 亚菊 Ajania pallasiana |
8 | 头状花序小,总苞宽2.5-4毫米,外层总苞片卵形、披针形,长1-2毫米,顶端无圆形蜡质附属物。 | 新疆亚菊 Ajania fastigiata |
9 | 叶不分裂或边缘锯齿或缺刻状锯齿或羽状全裂 (10) | |
9 | 叶二回羽状分裂或二回掌状或掌式羽状3-5分裂或三回羽状分裂或三回掌式羽状分裂。 (14) | |
10 | 叶不分裂或边缘锯齿或缺刻状锯齿。 (11) | |
10 | 叶羽状全裂;侧裂片1-4对,线形、披针形、长椭圆形或镰刀形。 (13) | |
11 | 叶全缘不分裂,线形。 | 柳叶亚菊 Ajania salicifolia |
11 | 叶边缘有缺刻状锯齿或锯齿。 (12) | |
12 | 叶下面被稠密厚绒毛,叶柄基部无叶耳。半灌木。 | 栎叶亚菊 Ajania quercifolia |
12 | 叶下面被稠密短柔毛,叶柄基部有分裂的叶耳;草本。 | 宽叶亚菊 Ajania latifolia |
13 | 侧裂片1-2对,线形,宽0.8-1.5毫米,边缘反卷。 | 异叶亚菊 Ajania variifolia |
13 | 侧裂片3-4对,长椭圆形、披针形、镰刀形或斜三角形,宽3-4.5毫米,边缘平。 | 分枝亚菊 Ajania ramosa |
14 | 小半灌木。木根系。 (15) | |
14 | 高大草本或铺散草本。须根系。 (18) | |
15 | 头状花序单生枝端。头状花序大,总苞直径7-10毫米。 | 单头亚菊 Ajania scharnhorstii |
15 | 头状花序在枝端排成伞房花序。 (16) | |
16 | 总苞片边缘紫黑色,全部花冠中部以上紫红色。 | 紫花亚菊 Ajania purpurea |
16 | 总苞片棕褐色,全部花冠黄色。 (17) | |
17 | 叶椭圆形、偏斜椭圆形,长1-2厘米,叶被稠密短绒毛。头状花序小,总苞宽4-5毫米。 | 西藏亚菊 Ajania tibetica |
17 | 叶全形半圆形、扁圆形,长0.3-0.5厘米,叶被稠密短柔毛。头状花序较大,总苞宽5-7毫米。 | 矮亚菊 Ajania trilobata |
18 | 叶二回羽状分裂或二回掌状或几掌状3-5分裂。 (19) | |
18 | 叶三回羽状分裂或三回掌式羽状分裂。 (26) | |
19 | 叶二回羽状分裂。 (20) | |
19 | 叶二回掌状或掌式羽状或几掌状3-5裂。总苞直径4-7毫米。 (24) | |
20 | 总苞片边缘膜质,内缘棕褐色或褐色,较宽,外缘白色膜质,较窄。 | 细叶亚菊 Ajania tenuifolia |
20 | 总苞片边缘全部褐色、棕褐色、黑褐色膜质,无内缘与外缘区别。 (21) | |
21 | 叶一、二回全部全裂。 (22) | |
21 | 一回为全裂、二回为浅裂、半裂或深裂。 (23) | |
22 | 叶两面灰白色,被稠密长或短柔毛。头状花序大,总苞直径宽5-6毫米。边缘雌花花冠顶端截形,花冠长1.2毫米,长为花柱之半。 | 短冠亚菊 Ajania brachyantha |
22 | 叶两面异色,上面绿色,几无毛;下面灰白色,被稠密短柔毛。头状花序小,总苞直径2.5-3毫米。边缘雌花花冠顶端明显3齿裂,花冠与花柱等长或几等长。 | 细裂亚菊 Ajania przewalskii |
23 | 一回侧裂片1-2对,如2对,则排列稀疏,裂距1厘米。末回裂片长椭圆形或镰刀形。 | 疏齿亚菊 Ajania remotipinna |
23 | 一回侧裂片2-4对,排列密集,裂距长5毫米。末回裂片椭圆形、披针形或斜三角形。 | 多花亚菊 Ajania myriantha |
24 | 末回裂片线形或狭线形。 | 丽江亚菊 Ajania adenantha |
24 | 末回裂片长椭圆形、椭圆形或倒披针状长椭圆形。 (25) | |
25 | 叶两面异色,上面绿色,几无毛;下面白色或灰白色,被稠密短柔毛。雌花花冠顶端3微齿。 | 短裂亚菊 Ajania breviloba |
25 | 叶两面同色或几同色,被等量稠密短柔毛。雌花花冠顶端3-4钝或深裂。 | 铺散亚菊 Ajania khartensis |
26 | 全株被稠密的厚绒毛,因而总苞片无清晰可见的轮廓。 | 密绒亚菊 Ajania sericea |
26 | 全株或仅叶上面稠密的短柔毛,总苞片外面被稀疏短柔毛或仅外层外面或外层基部被短柔毛,因而有清晰可见的黄褐色或黑褐色的膜质边缘部分和苞片本体。 (27) | |
27 | 叶大型,长3-3.5 厘米,宽2-3厘米,卵形或椭圆形,两面异色,上面绿色或淡绿色,无毛或几无毛。下面灰白毛,被稠密短柔毛。 | 多裂亚菊 Ajania tripinnatisecta |
27 | 叶小型,长宽1厘米,圆形或宽卵形,两面同色,灰白色,被等量的稠密短柔毛。 | 黄花亚菊 Ajania nubigena |
1 | Involucres straw-colored, glossy, 2-4 mm in diam.; phyllaries with colorless scarious margin | (2) |
1 + | Involucres not straw-colored, not glossy, 4-10 mm in diam.; phyllaries with dark brown or purple scarious margin | (15) |
2 (1) | Capitula 5-14, often regularly fascicled-corymbose at apices of branches; involucres cylindric or narrowly campanulate; phyllaries rigidly herbaceous, apex acute | (3) |
2 (1) + | Capitula many, corymbose or compound-corymbose at apices of stem and branches; involucres campanulate; phyllaries herbaceous, apex obtuse or rounded | (4) |
3 (2) | Leaf blade adaxially green, 2-pinnatisect. (23) | A. parviflora |
3 (2) + | Leaf blade gray-green, 3-parted or -lobed. (24) | A. trifida |
4 (2) | Perennial herbs | (5) |
4 (2) + | Small shrubs or subshrubs | (7) |
5 (4) | Involucres 6-7 mm in diam.; outer phyllaries narrowly elliptic, apices with rounded waxy appendage. (33) | A. pallasiana |
5 (4) + | Involucres 2.5-4 mm in diam.; outer phyllaries ovate or lanceolate, apices without rounded waxy appendage | (6) |
6 (5) | Stem weakly branched at base, often solitary, almost herbaceous; middle and lower stem leaf blades usually bipinnate with 5-7 lobes. (34) | A. fastigiata |
6 (5) + | Stem strongly branched at base, woody for much of length; middle and lower stem leaf blades usually twice ternately divided. (35) | A. gracilis |
7 (4) | Young stems glabrous; leaf blade abaxially green or pale green, glabrous or nearly so. (32) | A. nematoloba |
7 (4) + | Young stems hairy; leaf blade abaxially gray-white, appressed pubescent | (8) |
8 (7) | Leaf blade shallowly toothed to subentire, only occasionally lobed | (9) |
8 (7) + | Leaf blade pinnatisect, pinnatipartite, 2-pinnatisect, palmate-pinnately 3-5-divided, or twice ternately divided | (10) |
9 (8) | Leaf blade adaxially green or gray-green, glabrous or very sparsely pubescent; disk florets ca. 2.5 mm. (25) | A. potaninii |
9 (8) + | Leaf blade adaxially densely gray-white tomentose; disk florets ca. 4 mm. (26) | A. amphisericea |
10 (8) | Leaf blade pinnatisect or pinnatipartite | (11) |
10 (8) + | Leaf blade 2-pinnatisect, palmate-pinnately 3-5-divided, bipalmatisect, or twice ternately divided | (12) |
11 (10) | Leaf blade pinnatisect, adaxially gray-white, densely pubescent. (27) | A. truncata |
11 (10) + | Leaf blade pinnatipartite, adaxially green, glabrous. (28) | A. hypoleuca |
12 (10) | Leaf blade 2-pinnatisect. (29) | A. achilleoides |
12 (10) + | Leaf blade bipalmatisect, palmately-pinnately 3-5-divided, or twice ternately divided | (13) |
13 (12) | Leaf blade twice ternately divided. (35) | A. gracilis |
13 (12) + | Leaf blade bipalmately 3-5-sect or bipalmate, pinnately 3-5-sect | (14) |
14 (13) | Leaves with divided pseudostipules. (30) | A. nitida |
14 (13) + | Leaves with entire pseudostipules. (31) | A. fruticulosa |
15 (1) | Leaf blade undivided, 3-lobed, or simply pinnatisect, or pinnatipartite | (16) |
15 (1) + | Leaf blade more complexly divided: 2-pinnatisect, bipalmatisect, palmate-pinnately 3-7-divided, tripinnatisect, or tripinnate, pinnately divided | (21) |
16 (15) | Leaf blade undivided, margin entire, serrate, or incised-serrate | (17) |
16 (15) + | Leaf blade 3-lobed or pinnately divided | (19) |
17 (16) | Leaf blade linear, margin entire. (1) | A. salicifolia |
17 (16) + | Leaf blade narrowly elliptic, lanceolate, or obovate-oblong, or broadly oblanceolate, rarely linear, margin serrate or incised-serrate | (18) |
18 (17) | Subshrubs; leaves 5-10 × 0.3-4 cm, without pseudostipules, abaxially densely and thickly tomentose. (2) | A. quercifolia |
18 (17) + | Perennial herbs; leaves 3-4.5 × 1-2 cm, with divided pseudostipules, abaxially densely pubescent. (3) | A. latifolia |
19 (16) | Leaf blade 3-lobed. (6) | A. alabasica |
19 (16) + | Leaf blade pinnately divided | (20) |
20 (19) | Leaf blade with 1 or 2 pairs of lateral lobes; lobes linear. (4) | A. variifolia |
20 (19) + | Leaf blade with 3 or 4 pairs of lateral lobes; lobes narrowly elliptic, lanceolate, falcate, or obliquely triangular. (5) | A. ramosa |
21 (15) | Small subshrubs with thick, woody rootstocks; stems 4-25 cm | (22) |
21 (15) + | Erect or diffuse perennial herbs, with slender fibrous roots; stems often more than 25 cm | (25) |
22 (21) | Capitula solitary; involucres 7-10 mm in diam. (10) | A. scharnhorstii |
22 (21) + | Capitula in panicles; involucres 4-7(-8) mm in diam | (23) |
23 (22) | Phyllary margin dark purple; corolla apex purple-red. (8) | A. purpurea |
23 (22) + | Phyllary margin deep brown; florets uniformly yellow | (24) |
24 (23) | Leaf blade elliptic or obliquely elliptic in outline, 1-2 cm, both surfaces densely tomentose; involucre 4-6 mm in diam. (7) | A. tibetica |
24 (23) + | Leaf blade suborbicular in outline, 0.3-0.5 cm, both surfaces densely pubescent; involucre 5-7 mm in diam. (9) | A. trilobata |
25 (21) | Leaf blade tripinnatisect, subtripinnatisect, or palmate, tripinnately divided | (26) |
25 (21) + | Leaf blade 2-pinnatisect, bipalmatisect, or subpalmately 3-5-divided | (29) |
26 (25) | Whole plant densely and thickly tomentose; phyllaries inconspicuous. (19) | A. sericea |
26 (25) + | Whole plant pubescent or densely so; phyllaries conspicuous | (27) |
27 (26) | Leaf blade 3-3.5 × 2-3 cm, adaxially green or pale green, glabrous or nearly so, ultimate segments lanceolate or elliptic. (20) | A. tripinnatisecta |
27 (26) + | Leaf blade 1-2 × 1-1.5 cm, adaxially gray-white, densely pubescent, ultimate segments linear | (28) |
28 (27) | Stems thinly silvery gray tomentose; leaf blade ca. 2 × 1.5 cm. (21) | A. elegantula |
28 (27) + | Stems densely pubescent; leaf blade ca. 1 × 1 cm. (22) | A. nubigena |
29 (25) | Leaf blade bipalmatisect or palmately 3-5-divided | (30) |
29 (25) + | Leaf blade 2-pinnatisect | (31) |
30 (29) | Leaf blade adaxially green, subglabrous; involucres 4-5 mm in diam. (17) | A. breviloba |
30 (29) + | Leaf blade adaxially densely gray-white or white pubescent; involucres 6-10 mm in diam. (18) | A. khartensis |
31 (29) | Phyllary margin ± distinctly 2-colored: dark brown with a narrow but distinct colorless margin. (16) | A. tenuifolia |
31 (29) + | Phyllary margin uniformly brown or dark brown scarious | (32) |
32 (31) | Primary lateral segments of leaf blade pinnatilobed, pinnatifid, or pinnatipartite | (33) |
32 (31) + | Primary lateral segments of leaf blade or leaf blade pinnatisect | (34) |
33 (32) | Primary lateral segments of leaf blade 1- or 2-paired, if 2-paired then ca. 1 cm apart; ultimate lobes narrowly elliptic or falcate. (14) | A. remotipinna |
33 (32) + | Primary lateral segments of leaf blade 2-4-paired, ca. 5 mm at intervals; ultimate lobes elliptic, lanceolate, or obliquely triangular. (15) | A. myriantha |
34 (32) | Leaf blade adaxially gray-white, densely villous or pubescent; involucres 5-6 mm in diam.; corolla of marginal female florets with truncate apex. (11) | A. brachyantha |
34 (32) + | Leaf blade adaxially green, subglabrous; involucres 2.5-5 mm in diam.; corolla of marginal female florets with toothed apex | (35) |
35 (34) | Ultimate segments of leaf blade linear-lanceolate or narrowly elliptic; involucres ca. 2.5 mm in diam. (12) | A. przewalskii |
35 (34) + | Ultimate segments of leaf blade linear; involucres 4-5 mm in diam. (13) | A. adenantha |
多年生草本或亚灌木。叶互生,羽状分裂或掌式羽状分裂,极稀不分裂。头状花序小,多数或少数于茎、枝先端排成复伞房状或伞房状花序,稀单生于茎和枝顶,异型,边缘有5-15朵雌花,中央有多数两性花,全部小花均结实;总苞钟形或狭圆柱形;总苞片4-5层,草质,稀硬草质,先端和边缘为白色或褐色的膜质;花序托凸起或呈圆锥状,无托毛。雌花花冠管状或细管状,先端2-3齿或稀4-5齿;两性花花冠管状,冠檐稍扩大,先端5齿裂,全部小花花冠黄色,稀紫红色或带紫红色,外面具腺点;花柱分枝线形,先端平截;花药先端有披针形、尖或钝的附片,基部钝,无尾。瘦果有4-6条纵棱;冠毛无。约30种,主要分布于我国西北部和西南部,东北、华北和华中仅有少数种类;阿富汗北部、蒙古、俄罗斯和朝鲜北部也有少数种分布。云南产7种。
112.亚菊属——Ajania Poljak.
Poljak. in Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS, 17: 419. 1955; Tzvel. in Fl. URSS 26: 398. 1961. ——Chrysanthemum sect. Pyrethrum subsect. Dendranthema (DC.) Kitam. in Act. Phytotax. et Geobot. 4: 477. 1935.——Phaeostigma Muld. in Бот. Жур. 66(4):586. 1981.
多年生草本、小半灌木。叶互生,羽状或掌式羽状分裂,极少不裂的。头状花序小,异形,多数或少数在枝端或茎顶排列成复伞房花序、伞房花序,少有头状花序单生。边缘雌花少数,2-15个,细管状或管状,顶端2-3齿,少有4-5齿裂的。中央两性花多数,管状,自中部向上加宽,顶端5齿裂。全部小花结实,黄色,花冠外面有腺点,少有红紫色的。总苞钟状或狭圆柱状;总苞片4-5层,草质,少有硬草质,顶端及边缘白色或褐色膜质。花托突起或圆锥状突起,无托毛。花柱分枝线形,顶端截形,花药基部钝,无尾,上部有披针形的尖或钝附片。瘦果无冠毛,有4-6条脉肋。
本属约30种,主要分布我国除东南半壁以外的广大地区。蒙古、苏联及朝鲜北部和阿富汗北部也有少数种。
属模式种:Ajania pallasiana (Fisch. ex Bess.) Poljak.