| 1 |
Stamens long exerted; corolla ca. 0.8 cm
(35)
|
Dracocephalum stamineum
|
| 1 + |
Stamens included or only short exserted; corolla 1-4.2 cm.
|
(2)
|
| 2 (1) |
Dwarf cespitose plants, stems 3-15 cm; branches numerous, crowded; leaves 3-5 mm, circular-ovate, pinnati- or palmatipartite.
|
(3)
|
| 2 (1) + |
Stems erect or suberect, taller, unbranched or few branched; leaves more than 10 mm, simple to pinnatisect.
|
(5)
|
| 3 (2) |
Middle tooth of upper lip of calyx slightly wider than lateral teeth, upper lip teeth the same shape; leaves palmately 5-parted
(30)
|
Dracocephalum palmatoides
|
| 3 (2) + |
Middle tooth of upper lip of calyx 2.5-5 × wider than lateral teeth, broadly ovate, lateral teeth lanceolate to narrowly triangular; leaves pinnatifid.
|
(4)
|
| 4 (3) |
Middle tooth of upper lip of calyx ca. 2.5 × wider than lateral teeth
(31)
|
Dracocephalum origanoides
|
| 4 (3) + |
Middle tooth of upper lip of calyx 4-5 × wider than lateral teeth
(32)
|
Dracocephalum paulsenii
|
| 5 (2) |
Leaves pinnatisect, segments linear.
|
(6)
|
| 5 (2) + |
Leaves undivided.
|
(13)
|
| 6 (5) |
Calyx conspicuously 2-lipped; leaves sometimes 2-pinnatifid
(8)
|
Dracocephalum bipinnatum
|
| 6 (5) + |
Calyx of 5 subequal teeth, lanceolate to narrowly triangular; leaves clearly pinnatisect.
|
(7)
|
| 7 (6) |
Leaf sections 2- or 3-paired, crowded toward blade base, appearing palmatisect.
|
(8)
|
| 7 (6) + |
Leaf segments not crowded at blade base.
|
(10)
|
| 8 (7) |
Flowers 2.5-2.8 cm; leaf segments suberect
(5)
|
Dracocephalum forrestii
|
| 8 (7) + |
Flowers 3.5-4.2 cm; leaf segments oblique or subpatent.
|
(9)
|
| 9 (8) |
Calyx densely lanate-villous, teeth 1/4-1/3 as long as tube, teeth lanceolate-triangular, subulate- acuminate
(6)
|
Dracocephalum isabellae
|
| 9 (8) + |
Calyx slightly scabrid, teeth ca. as long as tube, lanceolate, acute
(7)
|
Dracocephalum hookeri
|
| 10 (7) |
Corolla less than 1.5 cm
(3)
|
Dracocephalum propinquum
|
| 10 (7) + |
Corolla more than 2 cm.
|
(11)
|
| 11 (10) |
Internodes 1-1.2 cm; leaves abaxially strigose on nerves
(1)
|
Dracocephalum taliense
|
| 11 (10) + |
Internodes 2-6 cm; leaves abaxially densely white pubescent.
|
(12)
|
| 12 (11) |
Upper floral leaves much shorter than calyx
(4)
|
Dracocephalum tanguticum
|
| 12 (11) + |
Upper floral leaves as long as to longer than calyx
(2)
|
Dracocephalum calophyllum
|
| 13 (5) |
Leaves entire.
|
(14)
|
| 13 (5) + |
Leaves serrate or dentate.
|
(17)
|
| 14 (13) |
Bracts 2- or 3-serrulate; corolla 1.4-1.8 cm; teeth of upper lip of calyx divided almost to base, middle tooth of upper lip ca. 2 × as wide as lateral teeth; anthers glabrous.
|
(15)
|
| 14 (13) + |
Bracts entire; corolla more than 1.8 cm; teeth of upper lip of calyx divided to nearly 1/2 its length, middle tooth of upper lip slightly wider than lateral teeth; anthers hairy.
|
(16)
|
| 15 (14) |
Leaves lanceolate to ovate-lanceolate, 1.5-3 × 4-8 mm, entire
(15)
|
Dracocephalum integrifolium
|
| 15 (14) + |
Leaves linear, 5-7 × 0.8-1.2 mm, 2- or 3-serrulate; teeth with apical spines to 3 mm
(14)
|
Dracocephalum fruticulosum
|
| 16 (14) |
Stems subglabrous; calyx minutely hairy toward base, subglabrous toward apex; corolla 3.3-4 cm
(34)
|
Dracocephalum argunense
|
| 16 (14) + |
Stems sparsely minute hairy toward base; calyx minutely hairy throughout; corolla 1.7-2.4 cm
(33)
|
Dracocephalum ruyschiana
|
| 17 (13) |
Calyx conspicuously 2-lipped.
|
(18)
|
| 17 (13) + |
Calyx inconspicuously 2-lipped.
|
(21)
|
| 18 (17) |
Calyx 2-lipped nearly to base
(11)
|
Dracocephalum breviflorum
|
| 18 (17) + |
Calyx 2-lipped to ca. 1/2 its length.
|
(19)
|
| 19 (18) |
Leaves ovate, base ± cordate, crenate; bracts 3-8-spinescent-serrate; corolla white
(9)
|
Dracocephalum heterophyllum
|
| 19 (18) + |
Leaves lanceolate or narrowly oblong, base cuneate to rounded, denticulate; bracts 1-3-spinescent-serrate; corolla blue-purple.
|
(20)
|
| 20 (19) |
Perennial herbs; leaves ± leathery, base cuneate, few spinescent-denticulate
(10)
|
Dracocephalum peregrinum
|
| 20 (19) + |
Annual herbs; leaves herbaceous, base rounded to broadly cuneate, densely dentate with teeth not spinescent or sometimes basal teeth spinescent
(12)
|
Dracocephalum moldavica
|
| 21 (17) |
Middle tooth of upper lip ± same shape as lateral teeth, nearly as wide as to wider than lateral teeth.
|
(22)
|
| 21 (17) + |
Middle tooth of upper lip of calyx wider than lateral teeth, circular-ovate; lateral teeth broadly lanceolate to narrowly triangular.
|
(25)
|
| 22 (21) |
Basal leaves absent; cauline leaves many paired, short petiolate, elliptic to ovate-elliptic, 5-6 mm, 1-3-denticulate; calyx teeth 5, subequal; bracts spinescent-serrate
(13)
|
Dracocephalum psammophilum
|
| 22 (21) + |
Basal leaves present.
|
(23)
|
| 23 (22) |
Cauline leaves many paired, broadly ovate; verticillasters in long spikes
(16)
|
Dracocephalum nutans
|
| 23 (22) + |
Cauline leaves 2-6-paired, ovate to triangular-ovate; verticillasters crowded in capitula.
|
(24)
|
| 24 (23) |
Calyx minutely papillate hirtellous, glandular
(19)
|
Dracocephalum rigidulum
|
| 24 (23) + |
Calyx pubescent to silky-villous, margin white ciliate
(20)
|
Dracocephalum imberbe
|
| 25 (21) |
Basal petiole much shorter than blade; blade ovate to broadly ovate, 0.6-2 cm, incised-dentate.
|
(26)
|
| 25 (21) + |
Basal petiole slightly shorter to longer than blade.
|
(27)
|
| 26 (25) |
Corolla yellowish white, middle lobe of lower lip not spotted
(17)
|
Dracocephalum nodulosum
|
| 26 (25) + |
Corolla purple, middle lobe of lower lip purple spotted
(18)
|
Dracocephalum hoboksarensis
|
| 27 (25) |
Corolla slightly exserted
(22)
|
Dracocephalum velutinum
|
| 27 (25) + |
Corolla 1.5-2 × as long as calyx.
|
(28)
|
| 28 (27) |
Middle tooth of upper lip of calyx subtruncate.
|
(29)
|
| 28 (27) + |
Middle tooth of upper lip of calyx rounded.
|
(30)
|
| 29 (28) |
Middle tooth of upper lip of calyx broadly elliptic, obtuse, short spinescent
(26)
|
Dracocephalum purdomii
|
| 29 (28) + |
Middle tooth of upper lip of calyx trapeziform, truncate, denticulate
(27)
|
Dracocephalum truncatum
|
| 30 (28) |
Leaves abaxially densely white pubescent
(21)
|
Dracocephalum wallichii
|
| 30 (28) + |
Leaves abaxially sparsely strigose.
|
(31)
|
| 31 (30) |
Petioles of middle cauline leaves 2-6 cm
(29)
|
Dracocephalum rupestre
|
| 31 (30) + |
Petioles of middle cauline leaves less than 1.2 cm or obsolete.
|
(32)
|
| 32 (31) |
Floral leaves coarsely dentate; corolla 3.3-4 cm
(28)
|
Dracocephalum grandiflorum
|
| 32 (31) + |
Floral leaves not coarsely dentate; corolla 2-3.5 cm.
|
(33)
|
| 33 (32) |
Leaves abaxially purplish, sparsely pubescent on veins or glabrous
(25)
|
Dracocephalum bullatum
|
| 33 (32) + |
Leaves abaxially densely puberulent, sparsely floccose-villous on veins.
|
(34)
|
| 34 (33) |
Bracts subcircular, subentire
(24)
|
Dracocephalum imbricatum
|
| 34 (33) + |
Bracts obovate to rhombic obovate, margin densely ciliate, 1-3-spinescent-serrate
(23)
|
Dracocephalum microflorum
|
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