学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
命名人 | (L.) Farwell. |
中文科名 | 金星蕨科 |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
中文属名 | 毛蕨属 |
学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
中文科名 | 金星蕨科 |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
中文属名 | 毛蕨属 |
学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
中文科名 | 金星蕨科 |
类型 | |
拉丁科名 | THELYPTERIDACEAE |
学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
中文科名 | 金星蕨科 |
类型 | |
拉丁科名 | Thelypteridaceae |
学名(拉丁名) | Cyclosorus parasiticus |
命名人 | (L.) Farw. |
发表年份 | 1931(年) |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
学名(拉丁名) | Cyclosorus parasiticus |
中文名 | 华南毛蕨 |
中文科名 | 金星蕨科 |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
中文属名 | 毛蕨属 |
学名(拉丁名) | Cyclosorus parasiticus |
发表年份 | 1931(年) |
拉丁属名 | Cyclosorus |
类型 | species |
拉丁科名 | Thelypteridaceae |
学名 | 状态 | 来源 | 命名人 |
---|---|---|---|
Polypodium parasiticum | Synonym | The Plant List | |
Aspidium parasiticum | Synonym | The Plant List | |
Cyclosorus parasiticus var. parasiticus | Synonym | The Plant List | |
Dryopteris parasitica | Synonym | The Plant List | |
Nephrodium parasiticum | Synonym | The Plant List |
植株高约50—70厘米。根状茎横卧至横走;叶近生至远生。叶柄长约30—40厘米,禾秆色,基部直径约3—4毫米,黑色,疏被披针形、暗棕色鳞片,向上疏被灰白色针状长毛或脱落后光滑。叶片卵状披针形,长约35—45厘米,宽约15—25厘米,一回羽状,顶端羽裂渐尖,基部的羽片不缩短或仅有1对略缩短。侧生羽片约10—15对,互生,条形,长约10—15厘米,宽约1—1.5厘米,顶端羽裂渐尖,基部截形,无柄,上侧较宽,边缘羽状半裂。 裂片约20—25对,长约4—5毫米,宽约4毫米,顶端钝圆,全缘。叶轴和羽轴禾秆色,密被灰白色针状长毛。叶草质,干后黄绿色,上面叶脉疏被针状毛,下面叶脉除密被灰白色针状长毛外还有黄红色腺体。每裂片上有小脉5—7对,基部1对连结,小脉单一,斜向上,伸达叶边。孢子囊群圆形,生于小脉的中部,在裂片中脉两侧各1行,通常靠近羽片中脉(羽轴)着生。囊群盖浅棕色,有白毛。
产于昆明、富宁、河口、绿春;生于林缘荒坡,海拔110—2000米。西藏、四川、贵州、广西、广东、海南、湖南、江西、浙江、福建、台湾也有。也分布于日本、韩国、越南、泰国、缅甸、马来西亚、印度尼西亚、菲律宾、印度、斯里兰卡。
Polypodium parasiticum Linnaeus, Sp. Pl. 2: 1090. 1793; Aspidium parasiticum (Linnaeus) Swartz; Christella parasitica (Linnaeus) H. Léveillé; Cyclosorus aureoglandulosus Ching & K. H. Shing; C. damingshanensis Ching ex K. H. Shing; C. excelsior Ching & K. H. Shing; C. hainanensis Ching; C. orientalis Ching ex K. H. Shing; C. rupicola Ching & K. H. Shing; C. xunwuensis Ching ex K. H. Shing & J. F. Cheng; C. yandongensis Ching & K. H. Shing; Dryopteris parasitica (Linnaeus) Kuntze; Nephrodium parasiticum (Linnaeus) Desvaux; Thelypteris parasitica (Linnaeus) Tardieu.
Plants (30-)50-70(-100) cm tall. Rhizomes shortly to long creeping, including stipe bases with dark brown lanceolate scales. Fronds approximate to distant; stipes (10-)20-30(-40) cm, stramineous; laminae (20-)30-50(-60) × (8-)15-25(-35) cm, bases not narrowed (sometimes slightly narrowed), apices caudate-acuminate; lateral pinnae 10-15(-20) pairs, proximal 1 or 2 pairs reflexed; middle pinnae lanceolate, (5-)10-15(-20) × (0.5-)1-1.5 cm, bases truncate, lobed 1/2-2/3 toward costae, apices long acuminate; segments 20-25 pairs, 3-4 × ca. 3 mm (basal acroscopic one longer, sometimes crenate), obtuse to subacute at apices; veinlets 5-8 pairs, simple (ca. 10 pairs on basal acroscopic segment, occasionally forked), proximal pair anastomosing, sometimes next vein running to sinus membrane. Laminae herbaceous, brownish green or yellowish green when dried, with thin acicular hairs throughout on both surfaces, and reddish orange glands throughout abaxially. Sori orbicular, medial; indusia densely hairy. Sporangia bearing reddish orange glands on stalks. Spores with wings. 2n = 72, 108, 144.
Semi-open places in thickets, roadsides; near sea level to 1900 m. Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang [N India, Indonesia, Japan, Korea, Laos, Myanmar, Nepal, Philippines, Sri Lanka, Thailand, Vietnam].
The type of Cyclosorus pauciserratus Ching & C. F. Zhang (Bull. Bot. Res., Harbin 3(3): 8. 1983) has no mature sori. Only one gathering has been made. This taxon might represent a young frond of C. parasiticus.
The types of Cyclosorus contractus Ching ex K. H. Shing (FRPS 4(1): 333. 1999) and C. parvilobus Ching ex K. H. Shing (FRPS 4(1): 338. 1999) both have the features intermediate between C. acuminatus and C. parasiticus. Each species has only one gathering (the type) with no sori collected. These two taxa might be hybrids.Cyclosorus ×intermedius W. C. Shieh & J. L. Tsai (J. Sci. Engin. 24: 8. 1987) is endemic to Taiwan. It is the putative hybrid between C. dentatus and C. parasiticus.The authors have not seen material of Cyclosorus parasiticus var. formosanus Ching (Bull. Fan Mem. Inst. Biol., Bot. 8: 205. 1938; Thelypteris parasitica var. formosana (Ching) C. F. Reed), described from Taiwan.The type of Cyclosorus brevipes Ching ex K. H. Shing (FRPS 4(1): 336. 1999) is partly bipinnate-pinnatifid with veins and glands similar to those of C. parasiticus. It has no sori and only one gathering has been collected. This taxon might be a hybrid.
33.华南毛蕨(中国主要植物图说,蕨类植物门)密毛毛蕨(台湾植物志,第二版)图版36:5-8
Cyclosorus parasiticus (L.) Farwell. in Amer. Midl. Naturalist 12: 259. 1931; H. Ito in Bot. Mag. Tokyo 51: 725. 1937. in Nakai et Honda, Nova Fl. Jap. No. 4: 176. 1939 et. Fil. Jap. Illustr. t. 356. 1944; Ching in Bull. Fan Mem. Inst. Biol. Bot. ser. 8: 201. 1938; in W. Y. Chun et al., Fl. Hainan 1: 126. 1964 et inY. L. Zhang et al. , Sporae Pterid Sin. 274. t. 61: 11-12. 1976; Tard.-Blot inLecomte, Not. Bot. Syst. 7: 75. 1938; Tard.-Blot et C. Chr., Fl. Indo-Chine 7(2): 381. 1941; Holtt., Fl. Mal. 2: Ferns Mal. 281 f. 162. 1945; 傅书遐, 中国主要植物图说 (蕨类植物门) 141. f. 185. 1957; Tagawa, Col. Ill. Jap. Pterid. 116. t. 45. f. 251. 1959; Fl. Fujian, 1: 157. 1982; Jarret, Ind. Fil. suppl. 4: 49. 1985; Shing in J. F. Cheng et G. F. Chu, Fl. Jiangxi 1: 209. f. 201. 1993; C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 172. f. 1176. 1993; Tsai et Shieh inFl. Taiwan 2ed 1: 174. photo 62. 1994; C. M. Zhang in W. T. Wang, Keys Vasc. Pl. Wuling. Mts. 42. 1995. ——Polypodium parasiticum L. Sp. Pl. 2: 1090. 1793; C. Chr. in Arkiv. Bot. Svenskka Vetensk. 9: 26. f. 4. 1910.——Aspidium parasiticum Sw. in Schrad. Journ. Bot. 1800, 2: 35. 1801; Christ, Farnkr. d. Erde 243. 1897 et in Warburg, Monsunia 1; 78. 1900. ——Nephrodium parasiticum Desv. Prodr. 260. 1827; Clarke in Trans Linn. Soc. 2, Bot. 1: 533. 1880. proparte; Diels in Engl. u. Prantl, Nat. Pflanzenfam. 1(4): 181. 1900; Cop., Polyp. Philip. 32., 1905. pro parte. ——Dryopteris parasitica O. Ktze., Rev. Gen. Pl. 2: 811. 1891: C. Chr., Ind. Fil. 282. 1906. pro parte; in Grad. Bull. Str. Settl. 4: 389. 1929. et Bull. Acad. Geogr. Bot. Mans 141. 1913; Merr., Enum. Hainan Pl. in Lingnan Sci Journ. 5: 9. 1927. pro parte; Ogata, Ic. Fl. Jap. 4: t. 147. 1931; Wu, Wong et Pong, Polyp. Yaoshan. in Bull. Dept. Biol. Sun Yatsen Univ. no. 3: 58. t. 20. 1932; Masamune, Fl. Hainan. 16. 1934.——Christella parasitica Lev., Fl. Kouy-tscheou 475. 1915; Holtt. in Kew Bull. 31(2): 309. 1976. pro parte et Fl. Males. ser 2, 1 (5): 559. f. 20 f. 1981; Kuo in Fl. Taiwan 1: 406. 1975; Edie, Ferns Hongk. 156. 1978; Jones et Clemesha, Austr. Ferns & Fern All. 116. f. 124. 1980. ——Thelypteris parasitica (L.) Tard.-Blot in Not. Syst. 7: 75. 193Fosb., Occ. Pap. B. P. Bishop Mus. 23: 30. 1962; K. Iwats. in Journ. Jap. Bot 38: 315. 1963. et in Mem. Coll. Sci. Univ. Kyoto B, 31(3): 172. 1965; Tagawa& K. Iwats. in Southeast Asia. St. 3(3): 79. 1965; 5: 65. 1967. et in Fl. Thailand 3(3): 424. 1988; Hatusima, Fl. Ryukyus 180. 1971; Kuo in Taiwania 30: 27. 1985; K. Iwarts., Ferns & Fern All. Jap. 219. pl. 142-5. 1992. ——Polypodium patens Sw., Prod. 133. 1788. ——Aspidium patens (Sw.) Sw. in Schrad. Journ.Bot. 1800(2): 34. 1801; Matsum. et Hay., Enum. Pl. Form. 581. 1906. ——Dryopteris patens O. Ktze. Rev. Gen. Pl. 2: 813. 1891; Hay., Ic. Pl. Form. 6: suppl. 188. 1916. ——Aspidium molle Benth, Fl. Hongk. 455. 1861; Christ, Farnkr. d. Erde 243. 1897. pro parte, non Sw. 1801. ——Nephrodium molle Hook.in Lond. Journ. Bot. 1: 494. 1842; Journ. Bot. 340. 1857. et Sp. Fil. 4: 67. 1862; Hook. et Bak., Syn. Fil. 293. 1867. pro parte; Bedd., Ferns S. Ind. t. 84, 1863. et Handb. Ferns Brit. Ind. 277. 1883; Franch. et Sav., Enum. Pl. Jap. 2:242. 1867; Dunn et Tutch., Fl. Kwangt. & Hongk. in Kew Bull. Add. ser. 10: 349. 1912. ——Nephrodium didymosorum Parish ex Bedd., Ferns Brit. Ind. t. 220. 1867. ——Nephrodium molle var. didymosorum Bedd., Handb. Ferns Brit. Ind. 279. 1883. ——Aspidium parasiticum var. didymosorum Christ in Warburg, Monsunia 1: 78. 1900. ——Dryopteris didymosora C. Chr., Ind. Fil. 262, 1906; v. A. v.R., Handb. Mal. Ferns 225. 1909; Bonap., Not. Pterid. pt. 7: 150. 1918. et pt. 14: 94. 1928. ——Cyclosorus didymosorus Nayar & Kaur, Comp. Bedd's Handb. 68. 1974. ——Aspidium procurrens Mett. in Ann. Mus. Bot. Lugd. Bat. 1: 231. 1964. ——Nephrodium procurrens (Mett.) Bak., Syn. Fil. 290. 1867; Bedd. Handb. Ferns Brit. Ind. suppl. 67. 1892. pro. parte.——Dryopteris procurrens (Mett.) Kuntze, Rev. Gen. Pl. 2: 813. 1891; v. A. v. R., Handb. Mal. Ferns211. 1908. pro parte.——Cyclosorus procurrens (Mett.) Cop., Fern Fl. Philip. 340. 1960. nom. tantum. ——Thelypteris procurrens (Mett.) Reed in Phytologia 17: 306. 1968. ——Dryopteris parasitica var. aureo-glandulosa Bonap., Not. Pterid. Pt.7: 149. 1918 et pt. 14: 94. 1923.
植株高达70厘米。根状茎横走,粗约4毫米,连同叶柄基部有深棕色披针形鳞片。叶近生;叶柄长达40厘米,粗约2毫米,深禾秆色,基部以上偶有一二柔毛;叶片长35厘米,长圆披针形,先端羽裂,尾状渐尖头,基部不变狭,二回羽裂;羽片12-16对,无柄,顶部略向上弯弓或斜展,中部以下的对生,相距2-3厘米,向上的互生,彼此接近,相距约1.5厘米,中部羽片长10-11厘米,中部宽1.2-1.4厘米,披针形,先端长渐尖,基部平截,略不对称,羽裂达1/2或稍深;裂片20-25对,斜展,彼此接近,基部上侧一片特长,约6-7毫米,其余的长4-5毫米,长圆形,钝头或急尖头,全缘。叶脉两面可见,侧脉斜上,单一,每裂片6-8对,(基部上侧裂片有9对,偶有二叉),基部一对出自主脉基部以上,其先端交接成一钝三角形网眼,并自交接点伸出一条外行小脉直达缺刻,第二对侧脉均伸达缺刻以上的叶边。叶草质,干后褐绿色,上面除沿叶脉有一二伏生的针状毛外,脉间疏生短糙毛,下面沿叶轴、羽轴及叶脉密生具一二分隔的针状毛,脉上并饰有橙红色腺体。孢子囊群圆形,生侧脉中部以上,每裂片(1-2) 4-6对;囊群盖小,膜质,棕色,上面密生柔毛,宿存。染色体2n=144。
产浙江南部及东南部、福建(崇安、福州)、台湾(台北、新竹、台中、南投、台南、高雄、台东、屏东)、广东(罗浮山、惠阳、怀集、信宜、鼎湖、大埔、徐闻、云浮)、海南(昌江、崖县)、湖南(宜章)、江西(井冈山、寻乌、定南)、重庆(缙云山)、广西(武鸣、大明山;龙州、百色、梧州)、云南东南部(河口)。生山谷密林下或溪边湿地,海拔90-1900米。日本、韩国、锡金、尼泊尔、缅甸、印度南部、斯里兰卡、越南、泰国、印度尼西亚(爪哇)、菲律宾均有分布。模式标本采自中国广东。