学名(拉丁名) | Aleuritopteris |
中文名 | 粉背蕨属 |
命名人 | Fee |
中文科名 | 中国蕨科 |
类型 | genus |
拉丁科名 | Sinopteridaceae |
学名(拉丁名) | Aleuritopteris |
中文名 | 粉背蕨属 |
中文科名 | 中国蕨科 |
类型 | genus |
拉丁科名 | Sinopteridaceae |
学名(拉丁名) | Aleuritopteris |
类型 | genus |
拉丁科名 | Pteridaceae |
学名(拉丁名) | Aleuritopteris |
中文名 | 粉背蕨属 |
类型 | genus |
拉丁科名 | Pteridaceae |
学名(拉丁名) | Aleuritopteris |
中文名 | 粉背蕨属 |
中文科名 | 中国蕨科 |
类型 | genus |
拉丁科名 | Sinopteridaceae |
学名(拉丁名) | Aleuritopteris |
发表年份 | 1850-1852 [1852](年) |
类型 | genus |
拉丁科名 | Pteridaceae |
1 | 叶片通常五角形,长宽几相等或长稍过于宽(由于顶部伸长);侧生羽片合生或少有下部1-3(4)对在往彼此以无翅的叶轴分开;裂片边缘有细密的钝齿(少有全缘);囊群盖连续不断,全缘或波状。 (2) | 银粉背蕨系 Ser. Argenteae Ching |
1 | 叶片长圆状披针形或长圆形,长2-4倍于宽;侧生羽片通常5-10对或更多,分离,彼此以无翅叶轴远分开,裂片全缘。囊群盖断裂(A.chrysophyila(HK.)Ching例外)彼此叠合,外缘具锯齿或撕裂。 (19) | 粉背蕨系 Ser. Farinosae |
2 | 叶片下面被白色、乳白色、乳黄色或硫黄色的蜡质粉末。 (3) | |
2 | 叶片下面不具蜡质粉末。 (17) | |
3 | 叶片下面连同叶柄、叶轴披卵圆披针形鳞片,幼时尤密,覆盖整个叶片。 | 毛叶粉背蕨 Aleuritopteris squamosa |
3 | 叶片下面无鳞片,或仅沿叶轴、羽轴具稀疏的不同鳞片。 (4) | |
4 | 叶片下面被雪白色粉末;叶柄的鳞片具锯齿;羽片分裂度细,叶片较小,长10厘米以下。 (5) | |
4 | 叶片下面被乳白色、乳黄色或硫磺色粉末;叶柄鳞片不具锯齿;植物体大,一般高15厘米以上。 (9) | |
5 | 叶片长过叶柄;叶柄及羽轴具鳞片及节状毛。 | 矮粉背蕨 Aleuritopteris pygmaea |
5 | 叶片短于叶柄,叶柄多少具鳞片而不具节状毛,羽轴光滑。 (6) | |
6 | 叶近二型,质薄;囊群盖狭,不连续。 | 莲座粉背蕨 Aleuritopteris rosulata |
6 | 叶一型,质厚;囊群盖宽,连续。 (7) | |
7 | 基部一对羽片与中央一片的分裂度相同,均为三向羽状深裂;叶柄粗壮,红棕色,每个孢子囊群只有1-2个孢子囊。 | 美丽粉背蕨 Aleuritopteris speciosa |
7 | 基部一对羽片与中央一片的分裂度不相同,叶柄纤细,褐黑色。 (8) | |
8 | 叶片长宽约在2-2.5厘米,裂片全缘,囊群盖宽几达中脉。 | 雪白粉背蕨 Aleuritopteris niphobola |
8 | 叶片一般长宽3-4.5厘米左右;裂片向顶部有锯齿;囊群盖较狭,远离中脉。 | 北京粉背蕨 Aleuritopteris niphobola var. pekingensis |
9 | 叶片下面的粉末为白色,密被粉红色腺体,因此下面呈粉红色。 | 丽江粉背蕨 Aleuritopteris likiangensis |
9 | 叶片下面的粉末为乳黄、硫磺或乳白色,不被红色腺体。 (10) | |
10 | 叶片下面的粉末为乳黄色或硫磺色。 (11) | |
10 | 叶片下面的粉末为乳白色或白色。 (12) | |
11 | 叶片下面被鲜明的硫磺色粉末;基部羽片的基部下侧一片小羽片全缘或近全缘。 | 硫磺粉背蕨 Aleuritopteris veitchii |
11 | 叶片下面被乳黄色粉末;基部羽片的基部下侧一片小羽片常为稀疏的深羽裂。 | 金爪粉背蕨 Aleuritopteris cremea |
12 | 下部第二对羽片不发达,远比相邻的上、下二对羽片短。 (13) | |
12 | 下部第二对羽片发达,比相邻的第三对羽片长,裂片狭,通常宽仅0.5厘米。 (14) | |
13 | 裂片宽披针形或几为三角形,彼此密接。 | 阔羽粉背蕨 Aleuritopteris tamburii |
13 | 裂片披针形,彼此远离。 | 雅砻粉背蕨 Aleuritopteris yalungensis |
14 | 下部第二对羽片基部心脏形,不下延或略下延,羽片彼此以无翅叶轴远分开。 (15) | |
14 | 下部第二对羽片基部下延成楔形,羽片彼此接近或以狭翅相连。 (16) | |
15 | 叶片五角形,长宽几相等;裂片较宽。 | 裂叶粉背蕨 Aleuritopteris argentea var. geraniifolia |
15 | 叶片卵圆形,长过于宽,裂片较狭。 | 假银粉背蕨 Aleuritopteris subargentea |
16 | 植株粗仕,高达30厘米以上,裂片宽,顶端羽片伸长成长尾状渐尖。 | 长尾粉背蕨 Aleuritopteris michelii |
16 | 植株高30厘米以下,顶端羽片不伸长成长尾状渐尖。 | 银粉背蕨 Aleuritopteris argentea |
17 | 植株较小,高15-20厘米(少有10厘米以下)叶片细裂,裂片线形。 | 陕西粉背蕨 Aleuritopteris shensiensis |
17 | 植株高大,高达20厘米以上,叶片粗裂,裂片长镰刀形。 (18) | |
18 | 侧生羽片5对,下部1-2对羽片羽裂(第二对仅下侧羽裂)裂片宽5-8毫米。 | 裸叶粉背蕨 Aleuritopteris duclouxii |
18 | 侧生羽片10对,下部3对羽片两侧均羽裂,裂片宽约3.5毫米。 | 多羽裸叶粉背蕨 Aleuritopteris nuda |
19 | 叶柄上密具暗棕色短腺毛;叶下面具雪白色粉末。 | 西畴粉背蕨 Aleuritopteris sichouensis |
19 | 叶柄上不具暗棕色短腺毛。 (20) | |
20 | 叶下面被鲜明的柠檬色粉末;囊群盖连续,全缘;叶柄鳞片线状披针形。 | 金粉背蕨 Aleuritopteris chrysophylla |
20 | 叶下面被白色、淡黄色或乳黄色粉末;囊群盖有锯齿或撕裂状;叶柄鳞片非线状披针形。 (21) | |
21 | 叶片下面、叶柄及叶轴被鳞片和伏生的节状毛。 (22) | |
21 | 叶片下面、叶柄及叶轴不被鳞片或伏生的节状毛(羽轴上偶有一、二鳞片)。 (24) | |
22 | 叶片下面被硫磺色粉末;沿羽轴和主脉有棕色钻形鳞片密生,羽片上面密被伏生的红棕色或灰色的节状毛。 | 棕毛粉背蕨 Aleuritopteris rufa |
22 | 叶片下面被白色或淡黄色的粉末;羽轴和主脉的鳞片披针形或卵状披针形,羽片上面无伏生的节状毛。 (23) | |
23 | 叶柄及叶轴上的鳞片宽披针形,中间黑色,两侧有阔而透明的白边,边缘具锯齿。 | 白边粉背蕨 Aleuritopteris albo-marginata |
23 | 叶柄上的鳞片披针形,具淡棕色狭边,边缘无锯齿。 | 多鳞粉背蕨 Aleuritopteris anceps |
24 | 叶片下面的粉末白色或雪白色;囊群盖边缘撕裂成睫毛状,叶片通常为卵圆披针形。 | 粉背蕨 Aleuritopteris pseudofarinosa |
24 | 叶下面粉末白色或乳黄色,囊群盖断裂,边缘撕裂状或有不规则的锯齿;叶片通常为长圆披针形。 (25) | |
25 | 囊群盖败育,孢子囊群裸露;叶片下面的粉末灰白色。 | 无盖粉背蕨 Aleuritopteris doniana |
25 | 囊群盖发达,叶下面的粉末白色或乳黄色。 (26) | |
26 | 叶片下面粉末为乳黄色或淡黄色,叶柄鳞片卵状披针形,边缘多少具腺体。 | 克氏粉背蕨 Aleuritopteris krameri |
26 | 叶片下面粉末白色或灰白色;叶柄鳞片边缘不具腺体。 (27) | |
27 | 植株较矮小,一般均在15厘米以下,密丛生;叶柄纤细。 | 阔盖粉背蕨 Aleuritopteris gresia |
27 | 植株较高大,一般均在20厘米以上;叶柄粗壮。 (28) | |
28 | 叶柄鳞片披针形,淡棕色。 | 台湾粉背蕨 Aleuritopteris formosana |
28 | 叶柄鳞片卵状披针形,棕红色。 | 狭盖粉背蕨 Aleuritopteris stenochlamys |
1 | Veins dark brown to black (sometimes partially obscured by farina), stout, prominently raised abaxially; sori with 1(or 2) large sporangium with broad annuli | (2) |
1 + | Veins green (sometimes obscured by farina), slender, not or only slightly raised abaxially; sori consisting of (1 or)2 to several small sporangia with narrow annuli | (3) |
2 (1) | Lamina 7-10 cm; medial pinnae pinnatifid; false indusia interrupted. (1) | A. grevilleoides |
2 (1) + | Lamina 3.5-8 cm; medial pinnae bipinnatifid; false indusia continuous. (2) | A. albofusca |
3 (1) | Lamina mostly pentagonal, nearly as wide as long; pinnae lobed to pinnatifid or only proximal 1-3 pairs of pinnae separated by wingless rachis; false indusia continuous, with entire or undulate margins | (4) |
3 (1) + | Lamina triangular-ovate, oblong-lanceolate, or oblong; pinnae usually 5-10 pairs or more, separated from each other by wingless rachis; false indusia usually interrupted, rarely continuous | (16) |
4 (3) | Lamina moderately to densely scaly on surface and veins abaxially. (3) | A. squamosa |
4 (3) + | Lamina lacking scales abaxially or scaly only on rachis and costae | (5) |
5 (4) | Lamina glabrous, lacking farina abaxially | (6) |
5 (4) + | Lamina with white or yellow farina abaxially (note that herbarium specimens dried with heat or treated with chemicals sometimes appear to lack farina and also rare hybrids involving farinose parental taxa sometimes do not produce farina) | (8) |
6 (5) | Pinnae curved upward; basal pinnae strongly inequilateral, acroscopic pinnules strongly reduced or lacking; lamina coarsely divided, ultimate segments falcate. (12) | A. duclouxii |
6 (5) + | Pinnae spreading or obliquely angled upward, moderately inequilateral, basal acroscopic pinnules developed, but shorter than basiscopic ones; lamina finely divided, ultimate segments relatively straight | (7) |
7 (6) | Second pair of pinnae (from lamina base) shorter than basal and third ones. (9) | A. tamburii |
7 (6) + | Second pair of pinnae (from lamina base) shorter than basal pair but longer than third ones. (11) | A. argentea |
8 (5) | Lamina usually less than 5 cm, with snow-white farina abaxially; sori consisting of (1 or)2 or 3 sporangia, discrete | (9) |
8 (5) + | Lamina usually more than 5 cm, with milky white or yellow farina abaxially; sori consisting of 3 to several sporangia, often confluent at maturity | (11) |
9 (8) | Laminae longer than stipes; stipes with scales and twisted hairs, these extending onto rachis and costae. (4) | A. pygmaea |
9 (8) + | Laminae shorter than stipes; stipes scaly only at base, not hairy; rachis and costae not hairy or scaly | (10) |
10 (9) | Lamina with basal pair of pinnae nearly equilateral, basal acroscopic segments lobed or pinnatifid, nearly as long as opposing basal basiscopic segments. (5) | A. speciosa |
10 (9) + | Lamina with basal pair of pinnae inequilateral, basal acroscopic segments simple, entire, much shorter than opposing basal basiscopic segments. (6) | A. niphobola |
11 (8) | Scales of rhizome concolorous, dark brown to black | (12) |
11 (8) + | Scales of rhizome bicolorous, with a brown to black central stripe and lighter margins | (13) |
12 (11) | Lamina with white farina abaxially and with relatively dense red glands (actually farina-producing glands with a red apical cell). (7) | A. likiangensis |
12 (11) + | Lamina with milky yellow farina abaxially, lacking red glands or some farina-producing glands with apical cell light brown, rarely somewhat reddish tinged. (8) | A. veitchii |
13 (11) | Lamina with light yellow to lemon-yellow farina abaxially. (12) | A. duclouxii |
13 (11) + | Lamina with white farina abaxially | (14) |
14 (13) | Second pair of pinnae (from lamina base) shorter than basal pair but longer than third ones. (11) | A. argentea |
14 (13) + | Second pair of pinnae (from lamina base) shorter than basal and third ones | (15) |
15 (14) | Medial pinnae rounded or bluntly pointed at tips; distal lobes of pinnae oblong, oblong-triangular, or nearly semicircular, with narrow V-shaped sinuses between lobes. (9) | A. tamburii |
15 (14) + | Medial pinnae attenuate at tips; distal lobes of pinnae linear to narrowly oblong, with broad U-shaped sinuses between lobes. (10) | A. yalungensis |
16 (3) | Lamina lacking farina at maturity (young fronds sometimes farinose) | (17) |
16 (3) + | Lamina with white or yellow farina abaxially | (19) |
17 (16) | False indusia continuous, margins undulate; lamina hairy abaxially. (28) | A. subvillosa |
17 (16) + | False indusia interrupted, margins laciniate; lamina glandular abaxially or glabrous except for inconspicuous hairs at pinna bases | (18) |
18 (17) | Fronds (15-)25-40 cm; stipe terete; lamina glabrous abaxially or occasionally sparsely and inconspicuously hairy at pinna bases. (26) | A. leptolepis |
18 (17) + | Fronds 7.5-15 cm; stipe grooved distally; lamina with brown glands along costae and veins abaxially. (27) | A. duthiei |
19 (16) | Stipe with short, dark brown glands. (13) | A. sichouensis |
19 (16) + | Stipe not glandular, but often scaly, occasionally also hairy | (20) |
20 (19) | Stipe, rachis, and costae scaly | (21) |
20 (19) + | Stipe scaly at base, sometimes up to rachis, but scales not extending to costae | (23) |
21 (20) | Scales on stipe broadly lanceolate, distinctly bicolorous, with conspicuous pale margins. (23) | A. albomarginata |
21 (20) + | Scales on stipe narrowly lanceolate, reddish brown, concolorous or mixed with occasional scales having narrow, lighter margins | (22) |
22 (21) | Lamina glabrous or rarely sparsely hairy adaxially; lamina widest at base. (24) | A. dubia |
22 (21) + | Lamina sparsely hairy and with sparse slender scales adaxially; lamina usually widest above base. (25) | A. rufa |
23 (20) | Lamina with milky yellow or golden-yellow farina abaxially | (24) |
23 (20) + | Lamina with white farina abaxially | (26) |
24 (23) | Stipe black, basalmost stipe scales black, slightly bicolorous, with very narrow, brown margins. (14) | A. ebenipes |
24 (23) + | Stipe usually reddish brown to dark brown, basalmost stipe scales concolorous, reddish brown | (25) |
25 (24) | Lamina with golden-yellow farina abaxially; stipe scales linear-lanceolate. (15) | A. chrysophylla |
25 (24) + | Lamina with milky yellow farina abaxially; stipe scales broadly lanceolate. (16) | A. krameri |
26 (23) | Stipe and rachis densely scaly. (17) | A. formosana |
26 (23) + | Stipe scaly proximally, rachis not scaly (occasionally sparsely scaly at base in A. anceps ) | (27) |
27 (26) | Rhizome scales brown, ovate-lanceolate, thin and translucent | (28) |
27 (26) + | Rhizome scales black, lanceolate, relatively thick and opaque | (29) |
28 (27) | Lamina hairy along costae and veins. (28) | A. subvillosa |
28 (27) + | Lamina lacking hairs. (29) | A. kuhnii |
29 (27) | Rhizome scales concolorous. (18) | A. gongshanensis |
29 (27) + | Rhizome scales bicolorous | (30) |
30 (29) | False indusia poorly developed to nearly absent. (19) | A. dealbata |
30 (29) + | False indusia well developed | (31) |
31 (30) | False indusia interrupted, with laciniate margins. (20) | A. anceps |
31 (30) + | False indusia continuous or sometimes interrupted, margins entire or undulate | (32) |
32 (31) | Lamina 2-6 cm, thinly herbaceous; false indusia poorly developed, narrow; stipe scales linear-lanceolate. (21) | A. rosulata |
32 (31) + | Lamina (3-)10-20 cm, leathery; false indusia well developed, broad; stipe scales broadly lanceolate. (22) | A. grisea |
常绿旱生中小型植物。根状茎短,直立或斜生,密生鳞片;鳞片披针形或卵状披针形,棕色或黑棕色,或中间黑棕色,边缘浅棕色或白色,全缘;叶簇生。叶柄圆柱形,棕色、栗黑色或乌木色,有光泽,光滑或疏生鳞片,具管状中柱。叶片五角形,三角形卵圆形或三角状长圆形,一至二回羽状。羽片无柄或近无柄;基部一对较大,三角形,一至二回羽裂。叶脉羽状,分离,纤细,通常不明显,伸达叶边。叶轴上面有纵沟,被面圆形。叶片纸质或草质,下面通常有白色、浅黄色或橙黄色的蜡质粉末,少数种类无粉末。孢子囊群生于叶边小脉先端,圆形,由2—10个孢子囊组成,彼此分离,但成熟时常向两侧扩展,彼此靠合而呈线形。囊群盖由反折的叶边特化而成,干膜质,棕色或灰棕色,稀为浅绿色,连续或不连续,边缘全缘、有锯齿或撕裂成睫毛状。孢子囊有短柄或近无柄;环带垂直,由15—18个细胞构成。孢子为球状四面形,极面对三角形或三角圆形,赤道面观长圆形,周壁光滑或有皱褶,或有颗粒状纹饰。染色体基数x = 15, 29 (30)。全世界30余种,分布于亚洲、非洲和热带美洲,以中国西部、南部和喜马拉雅山地区为本属的分布中心。中国有27种。云南有18种。常生于干旱地区含钙质的岩石或土壤上。
8.粉背蕨属——Aleuritopteris Fee
Fee Gen. Fil. 153, p1, 12. B. t. 1. 2. 1852; Ching in Hongk. Nat. 10: 194. 1941; Copel. Genera Filicum 1947 excl. Sinopteris
旱生常绿中小型植物。根状茎短而直立或斜升,密被鳞片;鳞片披针形,棕色或黑褐色或中间褐色,边缘浅棕色、淡白色,全缘。叶簇生,多数;叶柄和叶轴黑色,栗色或红棕色,有光泽,圆柱形,具管状中柱,无鳞片或稍具鳞片;叶片五角形,三角状卵圆形或三角状长圆形,2-3回羽状分裂,羽片无柄或几无柄,对生或近对生,基部一对较大,其基部下侧小羽片较大,伸长,下面通常具腺体,分泌黄色、白色或金黄色的腊质粉状物,偶有光滑;叶脉分离,羽状,纤细,通常不明显,达于叶边;叶轴上面有浅纵沟,下面圆。孢子囊群近边生,圆形,生于叶脉顶端,由2-10个孢子囊组成,彼此分离,成熟后常向两侧扩展成彼此接触;囊群盖干膜质,棕色、灰棕色或少有浅绿色,连续或不连续,边缘全缘或具锯齿或撕裂成睫毛状;孢子囊具短柄或几无柄,环带垂直,由15-18个细胞构成;孢子为球圆三角形,具三裂缝,周壁光滑或具皱褶,或具颗粒状纹饰。染色体x=29。
全世界30余种,中国有27种,为本属的现代分布中心。
模式种:Aleuritopteris, farinosa (Forsk.) Fee
全属可分下列两个系: