| 学名(拉丁名) | Cyrtomium |
| 中文名 | 贯众属 |
| 命名人 | Presl |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Cyrtomium |
| 中文名 | 贯众属 |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Cyrtomium |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Cyrtomium |
| 中文名 | 贯众属 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Cyrtomium |
| 中文名 | 贯众属 |
| 中文科名 | 鳞毛蕨科 |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 学名(拉丁名) | Cyrtomium |
| 发表年份 | 1836(年) |
| 类型 | genus |
| 拉丁科名 | Dryopteridaceae |
| 1 | 叶片为革质，羽片全缘无齿但有时边缘成波状。 (2) | |
| 1 | 叶片为纸质，羽片有齿或至少在近顶处有齿。 (14) | |
| 2 | 叶近似羽状复叶，顶生羽片羽裂渐尖状。 | 秦氏贯众 Cyrtomium chingianum |
| 2 | 叶为奇数羽状复叶，有分离的或大或小的顶生羽片，顶生羽片不为羽状分裂。 (3) | |
| 3 | 侧生羽片基部两侧多少内凹成心形。 (4) | |
| 3 | 侧生羽片基部一般不内凹，为圆楔形或宽楔形或近截形。 (9) | |
| 4 | 侧生羽片较小，长度一般不超过3厘米。 (5) | |
| 4 | 侧生羽片较大，长度在6厘米以上。 (6) | |
| 5 | 侧生羽片卵形，中部的长1-2厘米。 | 低头贯众 Cyrtomium nephrolepioides |
| 5 | 侧生羽片三角披针形，中部的长2-2.5厘米。 | 武陵贯众 Cyrtomium wulingense |
| 6 | 叶柄基部鳞片较宽，卵形，边缘流苏状 (7) | |
| 6 | 叶柄基部鳞片较狭，披针形，边缘有睫毛状齿。 (8) | |
| 7 | 羽片较小，长6-7厘米；小脉网眼较少，在主脉两侧各有2-3行。 | 邢氏贯众 Cyrtomium shingianum |
| 7 | 羽片较大，长12-14厘米；小脉网眼较多，在主脉两侧各有7-9行。 | 贵州贯众 Cyrtomium guizhouense |
| 8 | 通常仅具一个顶生羽片，有时其下部深裂成一对裂片，或至多成1对侧生羽片。 | 单叶贯众 Cyrtomium hemionitis |
| 8 | 叶为奇数羽状，侧生羽片1-5对。 | 厚叶贯众 Cyrtomium pachyphyllum |
| 9 | 羽片先端钝或圆形。 (10) | |
| 9 | 羽片先端变狭而伸长成渐尖状或尾状。 (13) | |
| 10 | 侧生羽片两侧及基部近对称。 (11) | |
| 10 | 侧生羽片两侧及基部不对称，基部偏斜近截形。 (12) | |
| 11 | 叶较大，羽片长3.5-4.5厘米。 | 惠水贯众 Cyrtomium grossum |
| 11 | 叶较小，羽片长1-2厘米。 | 世纬贯众 Cyrtomium tengii |
| 12 | 侧生羽片斜宽卵形或斜的扇形，长宽近相等。 | 新宁贯众 Cyrtomium sinningense |
| 12 | 侧生羽片三角卵形，长约为宽的2倍。 | 斜基贯众 Cyrtomium obliquum |
| 13 | 侧生羽片披针形。 | 披针贯众 Cyrtomium devexiscapulae |
| 13 | 侧生羽片偏斜的卵形或卵状披针形。 | 全缘贯众 Cyrtomium falcatum |
| 14 | 叶为羽状，先端羽裂渐尖状。 (15) | |
| 14 | 叶为奇数羽状复叶，顶端具2-3叉状或不分裂的顶生羽片。 (19) | |
| 15 | 羽片中脉两侧各具1行小脉网眼。 (16) | |
| 15 | 羽片中脉两侧各具2或3行小脉网眼。 (17) | |
| 16 | 羽片菱状卵形。 | 斜方贯众 Cyrtomium trapezoideum |
| 16 | 羽片镰状披针形。 | 单行贯众 Cyrtomium uniseriale |
| 17 | 羽片基部近对称，其上侧不具或仅有不明显的耳状凸。 | 尖羽贯众 Cyrtomium hookerianum |
| 17 | 羽片基部偏斜，其上侧有尖的耳状凸；羽片多少向上弯成镰状。 (18) | |
| 18 | 羽片上缘平直或向下成弯弓状；叶柄基部鳞片边缘具小尖齿。 | 镰羽贯众 Cyrtomium balansae |
| 18 | 羽片上缘多少向上隆起成弯弓状。 | 小羽贯众 Cyrtomium lonchitoides |
| 19 | 羽片有前倾的小齿或至少在顶部有齿，有时不明显近全缘。 (20) | |
| 19 | 羽片边缘有开张的尖齿。 (31) | |
| 20 | 侧生羽片基部近对称，宽楔形、圆楔形或楔形。 (21) | |
| 20 | 侧生羽片基部不对称，上下侧不同形。 (26) | |
| 21 | 侧生羽片卵形或基部1、2对为卵形。 (22) | |
| 21 | 侧生羽片为宽披针形、矩圆披针形、椭圆形或披针形等。 (24) | |
| 22 | 囊群盖边缘有齿。 | 齿盖贯众 Cyrtomium tukusicola |
| 22 | 囊群盖全缘无齿。 (23) | |
| 23 | 侧生羽片均为卵形，羽片背面有较多的披针形及纤毛状小鳞片 | 鳞毛贯众 Cyrtomium retrosopaleaceum |
| 23 | 基部1、2对羽片较大为卵形，其他为矩圆形，背面秃净或疏生小鳞片。 | 大叶贯众 Cyrtomium macrophyllum |
| 24 | 侧生羽片宽披针形、椭圆形或矩圆披针形，基部宽楔形至楔形；囊群着生处叶面平坦，也无明显凸起的囊托。 | 峨眉贯众 Cyrtomium omeiense |
| 24 | 侧生羽片披针形或线状披针形；囊群着生处叶背面下凹成小穴或有明显凸起的囊托。 (25) | |
| 25 | 囊群着生处叶背面常下凹成小穴，无明显凸起的囊托；囊群盖中央平坦或下凹，不凸起。 | 线羽贯众 Cyrtomium urophyllum |
| 25 | 囊群有明显凸起的囊托；囊群盖中央凸起成帽状。 | 台湾贯众 Cyrtomium taiwanense |
| 26 | 侧生羽片基部上侧楔形，下侧宽楔形。 | 福建贯众 Cyrtomium conforme |
| 26 | 侧生羽片基部上侧近截形，下侧楔形、宽楔形或圆楔形。 (27) | |
| 27 | 侧生羽片偏斜的卵形，少为宽披针形。 | 宽镰贯众 Cyrtomium latifalcatum |
| 27 | 侧生羽片披针形或矩圆披针形。 (28) | |
| 28 | 侧生羽片较小，一般长8厘米以下，基部上侧有时有耳状凸起。 (29) | |
| 28 | 侧生羽片较大，长8-12厘米，基部上侧有半圆形或三角形耳状凸起。 (30) | |
| 29 | 囊群盖全缘。 | 贯众 Cyrtomium fortunei |
| 29 | 囊群盖边缘流苏状或有小尖齿。 | 山东贯众 Cyrtomium shandongense |
| 30 | 侧生羽片较少，4-14对，彼此疏离。 | 阔羽贯众 Cyrtomium yamamotoi |
| 30 | 侧生羽片较多，20-22对，彼此密接。 | 密羽贯众 Cyrtomium confertifolium |
| 31 | 侧生羽片基部上侧有或大或小的耳状凸起。 (32) | |
| 31 | 侧生羽片基部上侧无耳状凸起。 (34) | |
| 32 | 侧生羽片基部上侧有长而尖的三角形耳状凸起。 | 刺齿贯众 Cyrtomium caryotideum |
| 32 | 侧生羽片基部上侧略有小而钝的耳状凸起，有时为半圆形。 (33) | |
| 33 | 囊群盖边缘有齿。 | 维西贯众 Cyrtomium neocaryotideum |
| 33 | 囊群盖全缘。 | 秦岭贯众 Cyrtomium tsinglingense |
| 34 | 侧生羽片较宽，卵形、矩圆卵形、矩圆披针形或宽披针形。 (35) | |
| 34 | 侧生羽片较狭，披针形。 (37) | |
| 35 | 羽片较薄，草质；囊群盖边缘微有齿。 | 膜叶贯众 Cyrtomium membranifolium |
| 35 | 羽片较厚，纸质；囊群盖全缘。 (36) | |
| 36 | 侧生羽片卵形。 | 尖齿贯众 Cyrtomium serratum |
| 36 | 侧生羽片宽披针形或矩圆披针形。 | 显脉贯众 Cyrtomium nervosum |
| 37 | 羽片基部近对称，楔形；囊群盖边缘有齿。 | 等基贯众 Cyrtomium aequibasis |
| 37 | 羽片基部不对称，圆楔形，上侧多少下延于羽柄；囊群盖全缘。 | 云南贯众 Cyrtomium yunnanense |
| 1 | Lamina leathery; pinna margins thickened, flat or sometimes repand | (2) |
| 1 + | Lamina papery, rarely subleathery or membranous; pinna margins flat, not thickened | (12) |
| 2 (1) | Terminal pinna pinnatifid and acuminate; lateral pinnae 18-22 pairs. (11) | C. chingianum |
| 2 (1) + | Terminal pinna simple, with 1 or 2 lobes or without lobes; lamina simple or with lateral pinnae 1-26 pairs | (3) |
| 3 (2) | Lateral pinnae cordate at base | (4) |
| 3 (2) + | Lateral pinnae rounded-cuneate, broadly cuneate, or truncate at base | (8) |
| 4 (3) | Lateral pinnae usually shorter than 3 cm, ovate or rarely deltoid-lanceolate. (1) | C. nephrolepioides |
| 4 (3) + | Lateral pinnae usually longer than 6 cm, oblong-ovate, oblong-lanceolate, deltoid-ovate, or ovate | (5) |
| 5 (4) | Stipe scales ovate, margins fimbriate | (6) |
| 5 (4) + | Stipe scales lanceolate, margins with ciliate teeth | (7) |
| 6 (5) | Lamina 18-24 cm; lateral pinnae 6-7 cm, acuminate; areoles 2 or 3 rows on each side of midrib; indusia entire. (2) | C. shingianum |
| 6 (5) + | Lamina ca. 42 cm; lateral pinnae 12-14 cm, acute; areoles 7-9 rows on each side of midrib; indusia toothed. (3) | C. guizhouense |
| 7 (5) | Lamina deltoid or ovate, simple or rarely with 2 or more lobes or 2 or more free lateral pinnae; venation palmate or ternate. (4) | C. hemionitis |
| 7 (5) + | Lamina oblong or triangular, imparipinnate, lateral pinnae 1-5 pairs; venation pinnate. (5) | C. pachyphyllum |
| 8 (3) | Pinnae acuminate or caudate at apex | (9) |
| 8 (3) + | Pinnae obtuse or rounded at apex | (10) |
| 9 (8) | Lateral pinnae lanceolate without acroscopic auricles; terminal pinna entire or lobed; indusia entire; stipe scales bicolorous, with dark brown centers and pale brown margins. (9) | C. devexiscapulae |
| 9 (8) + | Lateral pinnae obliquely ovate or ovate-lanceolate and with conspicuous acroscopic auricles; terminal pinna forked or trifurcate; indusia incised; stipe scales concolorous, pale brown. (10) | C. falcatum |
| 10 (8) | Two sides of midrib and base of lateral pinnae nearly equilateral; lateral pinnae 1-6 pairs. (6) | C. grossum |
| 10 (8) + | Two sides of midrib of lateral pinnae obviously inequilateral, base of lateral pinnae oblique and subtruncate; lateral pinnae 1-21 pairs, apex acute or obtuse | (11) |
| 11 (10) | Lamina 3-7 cm, acute; lateral pinnae 1-4 pairs, obliquely broadly ovate or obliquely flabellate, length subequal to width. (7) | C. sinningense |
| 11 (10) + | Lamina 13-35 cm, acuminate; lateral pinnae 12-21 pairs, ovate, ca. 2 ?as long as wide. (8) | C. obliquum |
| 12 (1) | Lateral pinnae with coarse teeth on margin | (13) |
| 12 (1) + | Lateral pinnae with only small teeth or sometimes subentire on margin | (19) |
| 13 (12) | Acroscopic base of lateral pinnae ?auriculate | (14) |
| 13 (12) + | Acroscopic base of lateral pinnae not auriculate | (15) |
| 14 (13) | Auricles of lateral pinnae small and blunt or semicircular; indusia entire on margin. (25) | C. tsinglingense |
| 14 (13) + | Auricles of lateral pinnae long and acute apically, deltoid; indusia irregularly dentate on margin. (26) | C. caryotideum |
| 15 (13) | Lateral pinnae lanceolate | (16) |
| 15 (13) + | Lateral pinnae ovate, oblong-ovate, oblong-lanceolate, or broadly lanceolate | (17) |
| 16 (15) | Stipe 22-36 cm; base of pinnae nearly equilateral, cuneate; indusia toothed on margin. (30) | C. aequibasis |
| 16 (15) + | Stipe 40-55 cm; base of pinnae inequilateral, rounded-cuneate and with acroscopic side ?decurrent; indusia entire on margin. (31) | C. yunnanense |
| 17 (15) | Leaves membranous; indusia denticulate on margin. (27) | C. membranifolium |
| 17 (15) + | Leaves papery; indusia entire on margin | (18) |
| 18 (17) | Lamina acute; lateral pinnae 8-13 pairs, ovate. (28) | C. serratum |
| 18 (17) + | Lamina obtuse; lateral pinnae 3-6 pairs, broadly lanceolate or oblong-lanceolate. (29) | C. anomophyllum |
| 19 (12) | Lateral pinnae inequilateral | (20) |
| 19 (12) + | Lateral pinnae nearly equilateral | (25) |
| 20 (19) | Acroscopic margins of lateral pinnae cuneate, basiscopic margins broadly cuneate. (24) | C. conforme |
| 20 (19) + | Acroscopic margins of lateral pinnae subtruncate, basiscopic margins cuneate, broadly cuneate, or rounded-cuneate | (21) |
| 21 (20) | Lateral pinnae obliquely ovate, rarely broadly lanceolate. (20) | C. latifalcatum |
| 21 (20) + | Lateral pinnae lanceolate or oblong-lanceolate | (22) |
| 22 (21) | Lateral pinnae usually shorter than 4 cm and narrower than 1.5 cm; acroscopic base obviously auriculate, auricles deltoid; basiscopic base cuneate. (12) | C. lonchitoides |
| 22 (21) + | Lateral pinnae up to 8-12 cm and 2.6 cm wide or wider; acroscopic base slightly or obviously auriculate; basiscopic base rounded or cuneate | (23) |
| 23 (22) | Lateral pinnae usually shorter than 8 cm; acroscopic base slightly auriculate, auricles blunt. (21) | C. fortunei |
| 23 (22) + | Lateral pinnae 8-12 cm; acroscopic base obviously auriculate, auricles semicircular or deltoid | (24) |
| 24 (23) | Lamina obtuse; lateral pinnae 4-14 pairs, distant; indusia incised or toothed. (22) | C. yamamotoi |
| 24 (23) + | Lamina acute; lateral pinnae 20-22 pairs, contiguous; indusia entire. (23) | C. confertifolium |
| 25 (19) | Lateral pinnae ovate or basal 1 or 2 pairs ovate | (26) |
| 25 (19) + | Lateral pinnae broadly lanceolate, oblong-lanceolate, or lanceolate | (28) |
| 26 (25) | Indusia toothed on margin. (15) | C. tukusicola |
| 26 (25) + | Indusia entire on margin | (27) |
| 27 (26) | Pinnae 2-6 pairs, 10-18 ?5-8 cm, ovate at base. (13) | C. macrophyllum |
| 27 (26) + | Pinnae more than 7 pairs, < 10 ?8 cm, rounded-cuneate to cuneate at base. (14) | C. muticum |
| 28 (25) | Lateral pinnae broadly lanceolate, elliptic, or oblong-lanceolate, base broadly cuneate to cuneate; both surfaces flat, without obvious apophyses (swellings on which sori are located) | (29) |
| 28 (25) + | Lateral pinnae lanceolate or linear-lanceolate, base cuneate; abaxial surface with pits containing sori or with obvious apophyses | (30) |
| 29 (28) | Lateral pinnae 4-9 pairs; basal pinnae not contracted; pinna margins entire or distal portion with small teeth; terminal pinna forked or trifurcate, 10-13 ?6-10 cm; indusia grayish brown in center, entire or toothed on margin; areoles 7 or 8 rows on each side of midrib. (17) | C. omeiense |
| 29 (28) + | Lateral pinnae 15-20 pairs; basal 2 pairs of pinnae contracted toward lamina base; pinna margins with small teeth; terminal pinna entire or lobed, 5-8 ?.8-2.5 cm; indusia dark brown in center, lacerate on margin; areoles 5 or 6 rows on each side of midrib. (16) | C. atropunctatum |
| 30 (28) | Lateral pinnae 8-13 pairs; terminal pinna obovate, ovate, or rhombic; sori contained in pits on abaxial surface of pinnae, without obvious apophyses; indusia flat or concave at middle. (18) | C. urophyllum |
| 30 (28) + | Lateral pinnae 6 or 7 pairs; terminal pinna oblong; sori on obvious apophyses; indusia raised and caplike at middle. (19) | C. taiwanianum |
土生，中小型常绿植物。根状茎短，直立或斜升，密被鳞片；鳞片大，质厚，卵状披针形至阔披针形，黑褐色，边缘有睫毛。叶簇生，一型。叶柄基部密被与根状茎上相同的鳞片，向上光滑。叶片卵状披针形，一回羽状。侧生羽片多对，卵形、卵状披针形或披针形，基部斜形，对称或近对称，基部上侧无耳状突起或有耳状突起，基部有短柄，边缘全缘或有锯齿，顶生羽片与侧生羽片同形。主脉明显，侧脉网状，在主脉两侧各有2-6行多边形的网眼，每网眼内具1-3条不分枝的内藏小脉。叶质地革质或厚纸质，两面无毛或沿叶轴和羽片中脉背面具纤维状鳞片。孢子囊群圆形，沿主脉两侧各有1-2行或多行，背生于网眼内的小脉上；囊群盖大，圆形，质厚，以中央一点盾状着生，宿存；孢子椭圆形，周壁具折皱，形成片状突起。染色体基数 x=41。约有50种，亚洲至非洲分布。中国有40种。云南有14种1变种。
11. 贯众属 Cyrtomium Presl
Presl, Tent. Pterid. 86. 1836; C. Chr. in Amer. Fern Journ. 20(2): 41. 1930; Ching in Bull. Chin. Bot. Soc. 2(2): 85. 1936.
陆生。根状茎短，直立或斜生，连同叶柄基部，密被鳞片。鳞片卵形或披针形，边缘有齿或流苏状。叶簇生，叶柄腹面有浅纵沟，嫩时密生鳞片，成长后渐脱落；叶片卵形或矩圆披针形少为三角形，奇数一回羽状或一回羽状，尖数仅具一枚顶生小叶（即单叶状）有时下部有1对裂片或羽片；侧生羽片多少上弯成镰状，其基部两侧近对称或不对称，有时上侧间或两侧有耳状凸起；主脉明显，侧脉羽状，小脉连结在主脉两侧成2至多行的网眼，网眼为或长或短的不规则的近似六角形，有内含小脉。叶纸质至革质，少有草质，背面疏生鳞片或秃净。孢子囊群圆形，背生于内含小脉上，在主脉两侧各1至多行；囊群盖圆形，盾状着生。
约40余种，主要分布在亚洲东部，以中国西南为中心，极少种类达印度南部和非洲东部。
本属的范围，特别是与热带美洲 Phanerophlebia 种类的关系，是有争议的问题。本志采用 C. Christensen (1930) ，秦仁昌（1936），邢公侠（1965）等人的概念。
KUN官网